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Transcript of NSave Nature to Survive - The Bioscan
NSave Nature to Survive
An International Quarterly Journal of Life Sciences
ISSN : 0973-7049Volume 13, Number 1: 2018
Published as an official organ by
NATIONAL ENVIRONMENTALISTS ASSOCIATION
P a g eP a g eP a g eP a g eP a g e
NSave Nature to Survive
CONTENTSCONTENTSCONTENTSCONTENTSCONTENTS
Website : www.thebioscan.com : www.neaindia.org
Cell : 94313-60645; 9572649448
Ph. : 0651-2244071
E-mails : [email protected]@gmail.com
For information regarding Association :SECRETARY,National Environmentalists Association,D-13, Sai Roofs, 1st Floor,H. H. Colony,Ranchi - 834002Jharkhand, India
Contact :For Editorial Information
Prof. M. P. SinhaVice ChancellorSido Kanhu Murmu UniversityDumka - 814 110Jharkhand, INDIA
NATIONAL ENVIRONMENTALISTSASSOCIATION
The National Environmentalists Associationis chartered in Ranchi as a nonprofit scientificand educational association of like mindedacademician, researchers, scientists from allover the nation for the furtherance and diffusionof knowledge of Life Sciences in general andEnvironmental Science in particular.
U.S .A . Of f i ce2827 Videre Dr.,Wilmington,DE 19808 We, USA
The association not only honours its membersbut also provides FELLOWSHIP tooutstanding contributors to the subject andthe society.
NAAS Rating : 5.26
A. RESEARCH PAPER
1. Evaluation of fungicides against leaf spot disease of apple (Malusdomestica cv. red delicious) caused by Alternaria mali RobertsU. S. Fais, V. Kumar and P. Vijimole
2. Chemical and sensory evaluation of flat bread supplemented withpearl millet and soy protein isolatesParamjot Kaur, Monika Sood and Julie D. Bandral
3. Genetic architecture of quality traits in bell pepper [Capsicumannum L. var. Grossum Sendt.]
Vibhuti Sharma and Sonia sood
4. Microbial diversity of moisture stress tolerant rhizobacteriaassociated with sorghum and allied weeds during sorghum cropproduction under drought condition
Kalindee S. Shinde and S. G. Borkar
5. Development and evaluation of extruded product of rice flourand apple pomaceMunazah Mehraj, H. R. Naik, Monica Reshi, S. A. Mir andA. Rouf
6. Sustainability through resource recycling, soil fertility and carbonsequestration from integrated farming systems in West coast, India
B. L. Manjunath, V. Paramesh, G. R. Mahajan, Bappa Das,K.Viswanatha Reddy, E. B. Chakurkar and N. P. Singh
7. Bio-efficacy of Tebuconazole 060 Fs (Raxil 060 FS) as seedtreatment against Karnal bunt, Loose smut and Flag smut of wheat
Bijender Kumar8. Assessment of genetic divergence among sugarcane genotypes
(Saccharum officinarum L.) based on molecular markersNaveen Arora, Lenika Kashyap, R. S . Gill and K. S Thind
9. Gamma ray induced viable mutants in soybean (Glycine max. (L.)Merrill)N. Verma, M. Chakraborty, K. Prasad, T. Izhar
10. Extent of heterosis over environments in rice hybrids using CMSsystemK. Parimala and CH. Surender Raju
11. Genetics of cotyledon colour in lentil (Lens culinaris Medik.)Yogesh Kumar, Jitendra Kumar and Sushil Kumar Chaturvedi
12. Catch efficiency of trawlers off Ratnagiri Coast of Maharashtra,IndiaB. R. Kharatmol, Latha Shenoy, V. V. Singh, A. T. Landgeand A. S. Mohite
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NSave Nature to Survive
PagePagePagePagePage
D I S C L A I M E R
The Publisher and Editors cannot beheld responsible for errors or anyconsequences arising from the use ofinformation in this journal; the viewsand opinions expressed do notnecessarily reflect those of thePublisher/ Association and Editors,neither does the publication ofadvertisements constitute anyendorsement by the Publisher /Association and Editors of the productsadvertised.
The Journal is CurrentlyAbstracted / Indexed in
© National Environmentalists Association
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13. Area under disease progress curve and influence abiotic factorson wilt of pomegranateR. Somu, R. K. Mesta, K. C. Kiran Kumar andChidanand P. Mansur
14. Exploitation of heterosis and inbreeding depression for yield,hybrid seed yield, nutritional and processing quality improvementin tomato (Solanum lycopersicum L.) under North IndianconditionsChandan Kumar and Surendra Prasad Singh
15. Correlation studies in some of the germplasm lines of kharifsorghumV. V. Kalpande, R. B. Ghorade, S. B. Thawari andY. G. Kedar
16. Genetic analysis of tropical maize (Zea mays L.) inbred linesunder heat stressAkula Dinesh, Ayyanagouda Patil, P. H. Zaidi, P. H.Kuchanur, M. T. Vinayan, K. Seetha Ram and AmareGouda
17. Heterosis studies for yield and yield components in greengram[Vigna radiata (L.) Wilczek]Anamika Nath and S. R. Maloo
18. Assessment of genetic diversity in germplasm collection of linseed(Linum usitatissimum L.)Chhaya Atri and Hitesh Kumar
19. Determination of selection criteria for grain yield in climateresilient small millet crop Kodo millet (Paspalum scrobiculatumL.)Abhinav Sao, Preeti Singh, Prafull Kumar and PraveenPanigrahi
20. Genetic assessment studies for yield and yield parameters indiverse genotypes of eggplant (Solanum melongena L.)Ashutosh Kumar, Anand Kumar Singh, Durga PrasadMoharana, Vaibhav Singh and Pushpendra Singh
21. Genetic variability and association analysis for yield and itscomponents in F
2 and F
3 segregating generations of the maize
(Zea mays L.) cross BM125 x BM135T. M. Deepu, S. K . Deshpande and B. R. Mani
22. Identification of best general combiners for yield and yield relatedtraits in maize (Zea mays L.) through pooled score techniquesB. C. Chandana and S. K. Deshpande
23. Studies on genetic variability, heritability and genetic advancein African marigold (Tagetes erecta L.)Jitendra Kumar Sahu, Gaurav Sharma, S. K. Nair and RamSingh
24. Correlation and path analysis in F2 generation of mungbean(Vigna radiata (L.) R. Wilczek)
A.Yusufzai Sana, M. S. Pithia, D. R. Mehta and Lata Raval
25. Combining ability and heterosis analysis for seed yield and it’scontributing traits in karingada [Citrulus lanatus (Thumb) Mansf.]
Satyendra Singh, S. D. Solanki, N. N. Prajapati and BharatChaudhary
26. Genetic variability and correlations in french bean for yieldimprovement
Savita, D. K. Singh and Ajay Tomer27. Influence of gamma rays on germination, survival and pollen
sterility in black gram (Vigna mungo L.) mutants
S. Anandhi Lavanya, C. Vanniarajan and J. Souframanien
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NSave Nature to Survive
ZONAL CO-ORDINATORS OF THEA S S O C I AT I O N
• Prof. N. BeheraSchool of Life Science,Sambalpur University
• Dr. Nirmal KumarISTAR, Vallabh Vidyanagar,Anand, Gujarat
• Dr. P. N. SudhaD. K. M. College for Women,Vellore
• Prof. S. P. S. DuttaDept. of Environmental Science,Jammu University, Jammu
• Dr. V. Salom Gnana ThangaDept. of Env. Scs.,University of Kerala, KariavattomTiruvananthapuram, Kerala
FEATURES OF ASSOCIATION
• Association is registered under80G of I.T.
• Prestigious fellowship of theAssociation (F. N. E. A.) toacademicians of the nation.
• Regular annual conference ofnational and international levelsorganized by the Association.
• Young Scientist and SeniorScientist award during theconference of the Association.
Publications of theAssociation
© National Environmentalists Association
An International Quarterly Journal of Life Sciences
An International Quarterly Journal of
Environmental Sciences
The Ecoscan
The Bioscan
www.theecoscan.inwww.thebioscan.in
Both the Journals areo n l i n e
Both the Journals areavailable on Google.com
Websites of the Journals are
PagePagePagePagePage28. Studies on effectiveness and efficiency of gamma rays in black
gram (Vigna mungo (L.) Hepper)
M. Dhasarrathan, S. Geetha, N. Meenakshiganesaan andD. Sassikumar
29. Hybridization of different gerbera (Gerbera jamesonii Bolus exHooker f.) lines for breeding novel traits
M. Aakanksha and Rajiv Kumar30. Screening of some long duration genotypes of pigeonpea [Cajanus
cajan (L.) Millisp.] against the infestation of pod fly[Melanagromyza obtusa (Malloch)]
Sabuj Ganguly, C. P. Srivastava and Sitanshu31. Estimation of genetic variability, heritability and genetic advance
in genotypes of maize (Zea mays L.)
Dan Singh Jakhar, Rajesh Singh, Vivek Kumar Ojha,Chandan Kumar and Aneesh Kumar Chandel
32. Estimation of heterosis for earliness and yield contributing traitsin cucumber (Cucumis sativus L.)
Manisha Thakur, Ramesh Kumar and Sandeep Kumar33. Genetic variability among genotypes and character association
in kharif potato (Solanum tuberosum L.) for different traits
Jyotshnarani Maharana, C. M. Panda and Praveen Jakhar34. Genotypic variation in pollen biology of jackfruit (Artocarpus
heterophyllus Lam.)
Aditi Guha Choudhury, Sonam Ongmu Bhutia, S. K. Sarkarand B. C. Das
35. SSR marker based parent selection of maize (Zea mays L.) inbredlines
Shashi Kumar, Rajesh Kumar, Ajay Kumar, NilmaniPrakash, Pankaj Kumar, Santosh Kumar, Gauttam Kumarand Vinay Kumar
36. Correlation and path coefficient analysis for yield and yieldcomponents in blackgram (Vigna mungo (L.) hepper)
Ch. Shalini and G. M. Lal37. Genetic variability, heritability and genetic advance in marigold
hybrids (Tagetes spp.)
V. P. Deepa and V. S. Patil38. Effectiveness and efficiency of gamma rays on sesame (Sesamum
indicum L.) genotypes
Sruba Saha and Amitava Paul
39. Genetic diversity analysis among mungbean genotypes based onrapd markers
L. Swathi, D. M. Reddy, K. H. P. Reddy and V. Sai Sruthi
40. Combining ability analysis for seed yield and its contributingtraits in linseed (Linum usitatissimum L.)
Ajeet Kumar, K. P. S. Tomar, V. Rathi, A. Kumar andN. K. Vasistha
41. Effect of pod related traits on shattering in M3 mutants ofmungbean (Vigna radiata (L.) wilczek)
N. Vairam
42. Genetic analysis of resistance to rice weevil Siltophilus oryzae L.in bread wheat
Ekhlaque Ahmad and J. P. Jaiswal43. Non-parametric methods for analyzing stability of chickpea (Cicer
arietinum L.) genotypes
A.Trivikrama Reddy, T. Lakshmi Pathy, A.Vijayabharathi andJayarame Gowda
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NSave Nature to Survive
THE BIOSCAN : SUBSCRIPTION RATESIndia SAARC Other(Rs.) Countries Countries
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44. Identification of hybrid rice (Oryza sativa L.) and its parental linesbased on morphological characters
Mampi Debbarma, Sharmila D. Deka and Debojit Sarma45. Molecular marker analysis of selected rice lines for aerobic traits
under water limited conditions
Kuldeep Kumar, Kanika Rani, Rahul Kumar Meena,Mahavir, Rajinder Kumar Jain and Sunita Jain
B. INSTRUCTION TO ATUTHORS
C. NATIONAL ENVIRONMENTALISTS ASSOCIATION: ANINTRODUCTION
231 - 236
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NSave Nature to Survive
13(1):207-214,2018www.thebioscan.com
207
COMBINING ABILITY ANALCOMBINING ABILITY ANALCOMBINING ABILITY ANALCOMBINING ABILITY ANALCOMBINING ABILITY ANALYSIS FOR SEED YIELD AND ITSYSIS FOR SEED YIELD AND ITSYSIS FOR SEED YIELD AND ITSYSIS FOR SEED YIELD AND ITSYSIS FOR SEED YIELD AND ITSCONTRIBUTING TRAITS IN LINSEED (CONTRIBUTING TRAITS IN LINSEED (CONTRIBUTING TRAITS IN LINSEED (CONTRIBUTING TRAITS IN LINSEED (CONTRIBUTING TRAITS IN LINSEED (LLLLLinuminuminuminuminum usitatissimumusitatissimumusitatissimumusitatissimumusitatissimum L.)L.)L.)L.)L.)
AJEET KUMAR1*, K. P. S. TOMAR1, V. RATHI1, A. KUMAR1 AND N. K. VASISTHA 2
1Department of Genetics and Plant Breeding,SVP University of Agriculture and Technology, Meerut - 250 110, INDIA2Department of Genetics and Plant Breeding, CCS University Meerut - 250 004, INDIAe-mail: [email protected]
INTRODUCTION
Linseed (Linum usitatissimum L.) is an ancient plant, which isalso known as flax. Linseed (2n = 30) is one of the first cropsdomesticated by man. Every part of the linseed plant is utilizedcommercially either directly or after processing. India is thethird largest producer in the world (Kiran et.al. 2012). In India,the crop is mainly cultivated in the states like Madhya Pradesh,Chhattisgarh, Uttar Pradesh, Maharashtra, Rajasthan, WestBengal, Karnataka, Orissa and Bihar. Chhattisgarh is one ofthe important linseed growing states of India, which account112.52 thousand hectare area and 34.20 thousand metrictons production (Deepak Gauraha et.al.2011). However, inthe world it covers 2270.35 thousand hectare area withproduction of 2238.94 thousand tons having productivity of986.16 kg per hectare, where as in India it covers 338thousand hectares area and a production of 147 thousandtons with the productivity of 434.91 kg per hectare,(Anonymous, 2013). Linseeds, particularly in their groundform are a great vegetarian source of the Omega 3 essential fattyacid, Alpha-Linolenic Acid (ALA). Linseed oil is an edible oil indemand as a nutritional supplement as a source of á-Linolenicacid (an omega-3 fatty acid) (Simopoulos 2002 and HassaneinM.S. et.al 2012). Combining ability is a powerful tool to selectgood combiners and thus selecting the appropriate parentallines for hybridization programme. In addition, the informationon nature of gene action will be helpful to develop efficientcrop improvement programme. General combining ability isdue to additive and additive × additive gene action and isfixable in nature while specific combining ability is due to
non-additive gene action which may be due to dominance orepistasis or both and is non-fixable. The presence of non-additive genetic variance is the primary justification forinitiating the hybrid breeding programme (Cockerham, 1961).Several workers like Singh et al. (1990), Khorgade et al. (1993)and Kumar and Paul (2015) have reported combining abilityon seed yield and its attributing traits in linseed.
There are several techniques for the evaluation of varieties orstrains in terms of their combining ability especially diallelmating system set excluding reciprocals. This technique wasdeveloped by Jinks and Hayman (1953). Keeping in view withthe above problem, the present investigation is taken up withthe following objectives: To study the general and specificcombining ability of parents and crosses for seed yield andyield contributing characters.
MATERIALS AND METHODS
The experiment field was conducted at Crop Research Centre,SardarVallabhbhai Patel University of Agriculture &Technology, Meerut (U.P.), during Rabi 2012-13. Theexperimental material comprised of ten parents namely, Sweta,Garima, Shekhar, Surbhi, Shubhra, Kiran, T-397, Neelam,Heera and Mukta and their all possible single cross was madeunder 10×10 diallel (Jinks 1953 and Hayman 1953) matingsystem set excluding reciprocals. The final trial was laid out inRandomized Block Design (RBD) with 45 F1’s and ten diversifiedparents during the year rabi 2013-14 in three replications.The observations were recorded on 5 randomly taken plantsper entry and per replication in case of parents and 45 crosses.
ABSTRACTA field experiment was conducted at Crop Research Centre (Chirodi), Sardar Vallabhbhai Patel University ofAgriculture & Technology, Meerut (U.P.), during Rabi 2012-13 and 2013-14 to estimate combining ability (GCAand SCA) of seed yield and its components of linseed. The experimental material comprised of ten parents andtheir 45s F2. The hybrids F2 were evaluated along with their parents in randomized complete block design withthree replications. It revealed highly significant variances for GCA and SCA for all the characters. Combiningability studies revealed that the lines Shekhar, Kiran, T-397 and Mukta, which were showed high generalcombining ability effects in desirable direction for seed yield and yield components. Moreover,. Among thespecific cross combinations, the crosses for seed yield per plant viz., Kiran×T-397, Shubhra×Hira, Neelam×Hira,T-397×Neelam and T-397×Hira exhibited good specific combining ability effects for seed yield and its contributingtraits.
KEYWORDSLinseedCombining abilitySeed yieldQuality characters
Received on :26.04.2017
Accepted on :21.05.2017
*Correspondingauthor
208
AJEET KUMAR et al.,
Data on the basis of randomly taken competitive plants wererecorded on viz., days to 50% flowering, days to maturity,plant height (cm), primary branches per plant, secondarybranches per plant, capsules per plant, seeds per capsule,biological yield per plant (g), seed yield per plant (g), harvestindex (%), 1000-seed weight (g) and oil content. Oil content ofeach genotype was determined by Soxhlet procedure (BISNo. 15:548 Part I 1964 and Official and Tentative Methods1973).
RESULTS AND DISCUSSION
Analysis of variance for combining abilityThe combining ability analysis under diallel mating approachwas done by the procedure advocated by Griffing (1956),which involves the study of covariances of full sibs andcovariances of half sibs to get the estimates of generalcombining ability and specific combining ability variancesand their effects. Analysis of variance for combining ability(Table 1) exhibited the variance due to general combiningability highly significant for all the characters and specificcombining ability variance was also observed highly significantfor all the traits, indicating involvement of both additive andnon-additive types of gene action in the inheritance of thesecharacters. Similar trend of involvement of both additive andnon-additive gene action has been earlier reported (Kalia,1972,Patil and Chopde, 1983, Khorgade et al., 1993), Popescu etal.,1995), Singh et al., 2008), Mohammadi et al., 2010, Yadavet al., 2013, Pali and Mehta, 2014).
The variance components for ó2s values were found higherthan estimates values ó2g for all the characters, suggestingmajor role of additive gene action in expression of these traits.Similar kind of additive gene action for various attributes wasreported earlier by Nie et al. (1991), Pillai et al. (1995), Awasthiand Rao (2005), Singh et al. (2008), Goral et al. (2008),Yadavet al. (2013) and Pali and Mehta (2014).
The proportion of ó2g / ó2s being less than unity for days to50% flowering, days to maturity, plant height, number ofprimary branches per plant, number of capsules per plant,number of secondary branches per plant, number of seedsper capsule, 1000 seed weight, biological yield per plant,harvest index, oil content and seed yield per plant suggestedthat involvement of non-additive kind of gene action for thesecharacters, which indicated that involvement of additive typeof gene action for this trait. The similar findings are inconformity with Mishra and Rai (1993), Popescu et al. (1995),Tiwari (1999), Awasthi and Rao (2005), Mohammadi et al.(2010) and Yadav et al. (2013).
The mean degree of dominance (ó2g / ó2s)½ was found greaterthan unity for all the characters, indicating the genetic controlof over dominance type of gene effect in them. These types offindings were also reported by Kaushal et al. (1974), Rai andDas (1975), Patil and Chopde (1983), Manfroni et al. (1989),Khorgade et al. (1993), Vishnu et al. (2005) and Yadav et al.(2013).
General combining ability effectsThe estimates of general combining ability effects and per seperformance of ten parents for all the 12 attributes are presented So
urce
of
D.F
.D
ays
toD
ays
toPl
ant
Num
ber
Num
ber
Num
ber
ofN
umbe
r of
1000
see
dB
iolo
gica
lH
arve
stO
ilSe
ed y
ield
vari
atio
n50
%m
atur
ityhe
ight
prim
ary
seco
ndar
yca
psul
es p
erse
eds
per
wei
ght
yiel
d pe
rin
dex
cont
ent
per
plan
tflo
wer
ing
(cm
)br
anch
esbr
anch
espl
ant
caps
ule
(g)
plan
t (g)
(%)
(%)
(g)
per
plan
tpe
r pl
ant
GC
A9
56.6
14**
32.6
93**
422.
033*
*2.
671*
*13
.120
**17
98.0
83**
0.65
8**
3.40
8**
3.41
1**
34.8
18**
5.19
5**
1.28
8**
SCA
4529
.149
**18
.676
**70
.893
**1.
685*
*5.
138*
*39
3.86
9**
0.90
1**
2.45
4**
5.03
8**
39.7
36**
2.61
6**
1.20
7**
Erro
r10
80.
784
1.22
20.
589
0.03
10.
209
5.59
90.
049
0.06
20.
094
2.22
40.
570
0.04
0
Tabl
e 1:
Ana
lysi
s of
var
ianc
e fo
r co
mbi
ning
abi
lity
of y
ield
and
its
com
pone
nts
in li
nsee
d(Li
num
usi
tati
ssim
umL.
)
Estim
ated
var
ianc
es d
ue to
σ2 g4.
653
2.62
335
.120
0.22
01.
076
149.
374
0.05
10.
279
0.27
62.
716
0.38
51.
104
σ2 s28
.365
17.4
5470
.304
1.65
44.
929
388.
270
0.85
22.
392
4.94
33.
512
2.04
51.
168
σ2 g/σ
2 s0.
164
0.15
00.
500
0.13
30.
218
0.38
50.
060
0.11
70.
056
0.07
20.
188
0.08
9(σ
2 s/σ
2 g)1/
22.
469
2.57
91.
414
2.74
22.
140
1.61
24.
087
2.92
84.
231
3.71
62.
305
3.35
1
209
COMBINING ABILITY ANALYSIS FOR SEED YIELD
Tabl
e 2:
Est
imat
es o
f GC
A e
ffec
ts a
nd p
er s
e pe
rfor
man
ce o
f par
ents
for
yiel
d an
d qu
ality
com
pone
nts
in li
nsee
d (L
inum
usi
tati
ssim
um L
.)
Cha
ract
ers
Day
s to
50%
flo
wer
ing
Day
s to
mat
urity
Plan
t hei
ght (
cm)
Num
ber
pri
mar
yN
umbe
r se
cond
ary
Num
ber
of c
apsu
les
bran
ches
per
pla
ntbr
anch
es p
er p
lant
per
plan
tPa
rent
sG
CA
Effe
ctM
ean
GC
A E
ffect
Mea
nG
CA
Effe
ctM
ean
GC
A E
ffect
Mea
nG
CA
Effe
ctM
ean
GC
A E
ffect
Mea
n
Swet
a-2
.638
**89
.407
-1.5
57**
140.
610
3.55
1**
65.3
170.
034
3.40
31.
276*
*16
.360
1.82
3**
124.
100
Gar
ima
-3.
325*
*81
.857
-3.4
45**
127.
873
5.13
8**
53.5
93-0
.230
4.77
70.
838*
*16
.873
5.62
3**
121.
287
Shek
har
0.5
3885
.097
-0.6
9113
2.92
01.
498*
*56
.597
0.2
586.
890
0.15
617
.310
1.91
8**
170.
233
Shub
hra
2.75
4**
96.7
53-0
.699
140.
103
2.5
16**
68.7
00-0
.073
5.73
3-0
.989
**12
.367
1.54
5*
126.
410
Kir
an -
0.40
886
.980
0.18
013
3.24
7-1
.886
**49
.967
0.0
315.
540
-0.0
8218
.100
5.00
7**
116.
100
Surb
hi0.
562
87.3
901.
193*
*13
7.65
75.
880*
*48
.953
-0.
127
4.37
70.
823*
*22
.867
10.4
09**
107.
300
T-39
7 -
1.91
0**
85.1
771.
555*
*14
5.01
0-4
.618
**56
.417
1.00
1**
7.86
31.
040*
*23
.290
26.7
06**
101.
467
Nee
lam
2.2
02**
91.9
531.
184*
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in (Table 2). The genotypes Garima (-3.325), Sweta (-2.638)and T-397 (-1.910) exhibited significant negative generalcombining ability effects and were found to be best generalcombiners for days to 50% flowering. The genotype Garima(-3.445) and Sweta (-1.557) exhibited significant negativegeneral combining ability effects and appeared to be bestgeneral combiners for early maturity. In the case of plant heightgenotypes T-397 (-4.618), Neelam (-3.475) and Kiran (-1.886)were found to be best general combiners for dwarfness andexhibited significant negative general combining ability effects.Out of ten parents, only three genotypes Hira (1.691), Mukta(1.526) and T-397 (1.001) showed significant positive generalcombining ability effects and appeared to be best generalcombiners for number of primary branches per plant. Thegenotypes Sweta (1.276), T-397 (1.040) Garima (0.838) andSurbhi (0.823) emerged as best general combiners for numberof secondary branches per plant which revealed significantand positive general combining ability effects. Among theparental lines, the genotypes T-397 (26.706), Mukta (15.928),Neelam (13.211), Surbhi (10.409), Hira (5.720), Garima(5.623), Kiran (5.007), Shekhar (1.918), Sweta (1.823) andShubhra (1.545) recorded significant and positive generalcombining ability effects for number of capsules per plant.Out of all parental lines, only three genotypes Surbhi (3.273),Shubhra (1.158) and Shekhar (1.105) revealed significant andpositive general combining ability effects and were found tobe best general combiners for number of seeds per capsules.The genotypes T-397 (3.745), Neelam (2.358), Shubhra (0.897)and Hira (0.412) exhibited significant positive generalcombining ability effects and emerged as best generalcombiners for 1000 seed weight. In the case of biologicalyield per plant the genotypes, Shekhar (2.128), Mukta (2.241),Kiran (1.554), T-397 (1.375) and Garima (1.031) exhibitedsignificant and positive general combining ability effects andappeared to be good general combiners for biological yieldper plant. Significant and positive general combining abilityeffects was observed for Kiran (3.164), T-397 (2.977), Mukta(2.294), Shekhar (1.894) and Sweta (1.117), which appearedas best general combiners for harvest index. The parents Mukta(5.537) and Sweta (1.278) revealed significant and positivegeneral combining ability effects and were emerged to be bestgeneral combiners for high oil content. Out of all parentallines, only four genotypes viz., T-397 (2.272), Shekhar (2.247),Mukta (1.291) and Kiran (1.080) observed significant andpositive general combining ability effects and were found tobe best general combiner for seed yield per plant. These resultsare in conformity with Thakur et al. (1988), Singh et al. (1990),Mishra and Rai (1993), Popescu et al. (1995), Tiwari (1999),Bhateria et al. (2002), Awasthi and Rao (2005), Goral etal.(2008), Yadav et al. (2013) and Pali and Mehta (2014).
Specific combining ability effectsThe specific combining ability effects and per se performanceof crosses is given in (Table-3). To confirm whether the crossesselected on the basis of specific combining ability effects werereally the best performance ones, the out of forty five crosscombinations, thirteen best cross combinations viz., Kiran×T-397 (11.180), Shubhra×Hira (10.967), Neelam×Hira(10.917), T-397×Hira (10.613), T-397×Neelam (10.296),Garima×Hira (9.515), Mukta×Hira (9.183), Sweta×Surbhi
210
Table 3: Estimation of SCA effects and per se performance of crosses for yield components in linseed (Linum usitatissimum L.)
Characters Days to 50% flowering Days to maturity Plant height(cm)Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Sweta x Garima 4.032** 84.240 0.267 135.593 4.886** 65.580Sweta x Shekhar 6.796** 90.867 -4.550** 136.807 -0.594 63.740Sweta x Shubhra -11.234** 75.053 -10.925** 135.593 -0.188 68.160Sweta x Kiran -12.771** 70.353 8.106** 136.807 -6.192** 58.753Sweta x Surbhi -3.888** 80.207 2.696* 135.593 4.002** 63.953Sweta x T-397 -2.190* 79.433 1.604 136.807 3.633** 64.847Sweta x Neelam 1.899* 87.633 -0.999 135.593 -2.984** 59.373Sweta x Mukta -2.778** 83.467 -6.124** 136.807 -8.114** 59.060Sweta x Hira 3.109** 86.153 -3.894** 135.593 13.683** 93.600Garima x Shekhar 2.616** 86.000 -1.369 136.807 -0.079 55.567Garima x Shubhra -1.067 84.533 -2.421* 135.593 -0.393 59.267Garima x Kiran -16.471** 65.967 1.800 136.807 -3.057** 53.200Garima x Surbhi 4.246** 87.653 -0.530 135.593 -0.730 50.533Garima x T-397 -3.703** 77.233 0.945 136.807 3.125** 55.650Garima x Neelam 2.853** 87.900 3.446** 135.593 5.172** 58.840Garima x Mukta 2.442** 88.000 3.820** 136.807 -0.269 58.217Garima x Hira 0.382 82.740 -1.856 135.593 -11.831** 59.397Shekhar x Shubhra -1.263 88.200 2.622* 136.807 3.844** 67.143Shekhar x Kiran 2.486** 88.787 -1.487 135.593 -3.080** 56.817Shekhar x Surbhi -2.837** 84.433 4.279** 136.807 -0.660 54.243Shekhar x T-397 -1.849* 82.950 -0.649 135.593 -0.405 55.760Shekhar x Neelam -0.377 88.533 -4.738** 136.807 -0.682 56.627Shekhar x Mukta 2.319** 91.740 7.806** 135.593 -4.759** 57.367Shekhar x Hira -3.588** 82.633 3.113** 136.807 11.792** 86.660Shubhra x Kiran 8.483** 97.000 -1.275 135.593 -6.361** 57.550Shubhra x Surbhi 0.097 89.583 0.035 136.807 9.476** 68.393Shubhra x T-397 -0.832 86.183 -0.730 135.593 1.854 62.033Shubhra x Neelam -0.573 90.553 -4.313** 136.807 7.218** 68.540Shubhra x Mukta 0.279 91.917 3.765** 135.593 -7.336** 58.803Shubhra x Hira -4.037** 84.400 4.869** 136.807 -10.889** 67.993Kiran x Surbhi 6.326** 92.650 -0.622 135.593 -6.814** 48.700Kiran x T-397 2.447** 86.300 0.850 136.807 -0.577 56.200Kiran x Neelam 3.302** 91.267 -2.586** 135.593 0.600 58.520Kiran x Mukta 4.458** 92.933 0.495 136.807 28.603** 91.340Kiran x Hira -1.508 83.767 3.566** 142.533 17.961** 93.440Surbhi x T-397 1.477 86.300 -2.363** 137.200 5.987** 57.770Surbhi x Neelam -0.207 88.727 2.034** 141.227 3.444** 56.370Surbhi x Mukta -3.028** 86.417 2.985** 141.300 -1.199 56.543Surbhi x Hira -2.378** 83.867 -5.424** 134.557 -10.372** 60.113T-397 x Neelam -0.829 85.633 -1.554 138.000 -3.065** 51.123T-397 x Mukta 1.936* 88.910 0.330 139.007 -3.318** 55.687T-397 x Hira -2.106* 81.667 -7.606** 132.737 -13.978** 57.770Neelam x Mukta -2.218** 88.867 3.221** 141.527 -1.508 58.640Neelam x Hira -6.608** 81.277 5.068** 145.040 -13.274** 59.617Mukta x Hira -6.978** 81.417 -0.317 138.777 12.222** 89.930S.E.(Sij) 0.644 0.052 1.425SE(Sij-Sik) 1.417 0.016 0.095
Characters Number Primary branches Number secondary branches Number of capsulesper plant per plant per plant
Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Sweta x Garima -1.088 3.733 -0.326 18.400 17.101** 123.427Sweta x Shekhar 0.124 5.433 0.633 18.667 16.089** 120.733Sweta x Shubhra 1.489 6.467 -0.489 16.400 15.584** 152.033Sweta x Kiran 1.585** 6.667 0.837 18.633 10.902** 140.800Sweta x Surbhi 0.376 5.300 1.126 19.827 15.005** 139.500Sweta x T-397 1.481 7.533 2.722** 21.640 17.901** 126.100Sweta x Neelam 1.374 6.747 0.879 18.467 -43.216** 104.900Sweta x Mukta -0.385 4.140 -1.744 15.387 21.201** 172.033Sweta x Hira -1.593 2.767 1.943 17.797 13.776** 154.400
Table 3: Cont......
AJEET KUMAR et al.,
211
* Significant at 5% probability level; ** Significant at 1% probability level
Characters Number Primary branches Number secondary branches Number of capsulesper plant per plant per plant
Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Garima x Shekhar -2.611** 2.433 1.171 18.767 18.442** 162.710Garima x Shubhra 0.520 5.233 -0.884 15.567 34.075** 177.970Garima x Kiran -0.117 4.700 0.529 17.887 40.940** 178.283Garima x Surbhi 3.507** 8.167 2.937** 21.200 -10.797** 121.143Garima x T-397 -0.221 5.567 2.113*** 16.367 37.882** 153.527Garima x Neelam -2.542** 3.567 1.084 18.233 7.855** 147.707Garima x Mukta -0.028 4.233 0.767 17.460 18.494** 139.783Garima x Hira 3.138** 5.233 -0.366 15.050 23.720** 124.350Shekhar x Shubhra -0.968 4.233 -1.552 14.217 -11.956** 128.233Shekhar x Kiran -0.972 4.333 -0.492 16.183 -16.781** 116.857Shekhar x Surbhi 2.086** 7.233 0.053 17.633 0.059 102.177Shekhar x T-397 -1.209 5.067 -4.148** 13.650 0.906 101.033Shekhar x Neelam 0.703 6.300 0.667 17.133 -1.506 150.350Shekhar x Mukta -0.516 4.233 3.723** 17.733 0.171 157.743Shekhar x Hira 0.650 5.233 1.150 15.883 0.342 146.707Shubhra x Kiran -1.440 3.533 1.969 17.500 -16.605** 116.660Shubhra x Surbhi -1.483 3.333 -0.135 16.300 -14.229** 113.633Shubhra x T-397 -0.878 5.067 1.247 17.900 -0.230 108.337Shubhra x Neelam -1.032 4.233 2.911** 18.233 13.423** 164.907Shubhra x Mukta 2.049** 6.467 0.501 15.367 6.270** 160.470Shubhra x Hira 0.015 4.267 0.945* 14.533 0.481 147.473Kiran x Surbhi 2.386** 3.533 5.609** 11.733 -0.211 112.100Kiran x T-397 -0.548 5.500 4.360** 13.200 -1.948 86.067Kiran x Neelam -0.836 4.533 1.371 17.600 17.818** 162.750Kiran x Mukta 0.612 5.133 1.894 17.667 11.619** 159.267Kiran x Hira 2.177** 6.533 0.538 15.033 1.493 138.933Surbhi x T-397 -0.357 5.533 2.132** 16.333 1.621 101.233Surbhi x Neelam -0.578 4.633 -3.501** 15.633 31.438** 170.967Surbhi x Mukta -0.830 3.533 -1.977 14.700 14.355** 156.600Surbhi x Hira -0.565 3.633 -2.000** 13.400 15.096** 147.133T-397 x Neelam -0.140 6.200 0.715 18.067 -0.966 99.267T-397 x Mukta 0.441 5.933 0.138 17.033 -0.682 105.267T-397 x Hira -0.260 5.067 -1.285 14.333 15.975** 99.767Neelam x Mukta -0.280 4.533 2.303** 17.867 0.234 171.100Neelam x Hira -0.848 3.800 -1.987 12.300 0.508 158.167Mukta x Hira 3.400** 4.200 -0.597 13.233 -0.041 155.333S.E.(Sij) 0.327 0.849 1.393SE(Sij-Sik) 0.481 0.249 1.457
Table 3: Cont.....
Characters Number of seeds per capsule 1000 seed weight (g) Biological yield per plant (g)Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Sweta x Garima 0.478 9.433 0.753 8.030 -0.852 14.110Sweta x Shekhar -0.922 7.100 -10.800** 7.643 1.897 15.700Sweta x Shubhra 0.442 8.200 0.382 8.213 -0.339 13.113Sweta x Kiran -0.470 7.400 -0.526 8.200 0.971 15.457Sweta x Surbhi -1.690 6.500 -1.537 6.877 2.197** 16.123Sweta x T-397 0.318 8.067 0.148 8.337 -0.163 13.393Sweta x Neelum -0.080 7.533 -2.043** 6.533 -1.18 12.733Sweta x Mukta -1.147 6.467 2.359** 8.650 3.027** 17.200Sweta x Hira 0.337 8.600 2.587** 8.933 1.144 15.110Garima x Shekhar 1.022 8.867 10.683** 8.153 -10.909** 14.787Garima x Shubhra 3.885** 8.467 10.041 8.900 12.101** 13.243Garima x Kiran -0.126 7.567 11.366** 8.120 -0.311 16.067Garima x Surbhi 0.321 8.333 0.535* 8.977 1.975 17.793Garima x T-397 -0.238 7.333 -0.087 8.130 -1.049 14.400Garima x Neelam 5.636** 6.800 0.323 8.927 -0.630 15.183Garima x Mukta 10.497** 7.933 -0.658 8.660 -3.542** 12.523Garima x Hira -0.553 7.533 0.326 9.700 -1.852 14.007Shekhar x Shubhra -4.181** 6.467 -0.169 8.857 -0.719 13.467
Table 3: Cont.....
COMBINING ABILITY ANALYSIS FOR SEED YIELD
212
* Significant at 5% probability level; ** Significant at 1% probability level
Characters Number of seeds per capsule 1000 seed weight (g) Biological yield per plant (g)Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Shekhar x Kiran 7.042** 8.800 4.123** 8.043 2.188** 17.407Shekhar x Surbhi 1.255 9.333 -0.271 7.337 0.787 15.447Shekhar x T-397 -0.204 7.433 -0.850 6.533 0.034 14.323Shekhar x Neelam -1.102 6.400 -0.517 7.253 2.121** 12.533Shekhar x Mukta 0.331 7.833 -1.601 6.883 2.912** 11.993Shekhar x Hira -1.552 6.600 1.070 9.610 1.634** 16.333Shubhra x Kiran -1.662 5.833 2.509** 8.817 -0.324 14.543Shubhra x Surbhi 0.218 8.033 2.397** 8.393 -1.412 12.897Shubhra x T-397 8.059** 8.433 1.835 8.607 1.682 15.620Shubhra x Neelam -0.039 7.200 -1.545 7.613 -0.647 13.657Shubhra x Mukta 9.028** 8.267 -0.296 8.577 2.799** 17.353Shubhra x Hira -0.822 7.067 -0.242 8.687 -1.465 12.883Kiran x Surbhi -0.526 7.400 -0.774 7.117 3.485** 18.827Kiran x T-397 -1.018 6.467 -1.103** 6.563 -1.178 13.793Kiran x Neelam 0.850 8.200 0.417 8.470 -2.207** 13.130Kiran x Mukta 0.117 7.467 -2.084** 6.683 -3.108** 12.480Kiran x Hira 0.033 8.033 11.396** 7.427 -1.515** 13.867Surbhi x T-397 -1.338 6.467 -0.761 6.593 0.588* 15.000Surbhi x Neelam 10.797** 8.467 -0.055 7.687 3.111** 11.667Surbhi x Mukta 11.397** 9.067 10.782 7.673 -0.219 14.810Surbhi x Hira -0.553 7.767 -0.614 7.897 -1.619 13.203T-397 x Neelam -0.529 6.700 1.533 9.050 -0.338 14.070T-397 x Mukta -1.096 6.133 0.339 8.570 -0.065 14.593T-397 x Hira 0.454 8.333 -1.063 7.223 -0.185 14.267Neelam x Mukta 0.439 7.533 3.549** 8.167 13.391** 11.633Neelam x Hira 0.089 7.833 -1.240 7.433 3.073** 17.890Mukta x Hira 0.223 7.967 2.079** 8.467 0.695 15.763S.E.(Sij) 0.413 0.461 0.570SE(Sij-Sik) 0.607 .678 0.838
Table 3: Cont.....
Characters Harvest index (%) Oil content (%) Seed yield per plant (g)Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Sweta x Garima 2.125** 56.910 -0.187 41.617 -0.165 8.023Sweta x Shekhar 4.945** 61.640 0.490 42.333 5.847** 9.677Sweta x Shubhra 3.385* 56.143 1.374 43.157 0.282 7.333Sweta x Kiran 4.237** 57.873 -0.696 39.987 6.280** 8.943Sweta x Surbhi -0.829 54.203 -1.845 39.833 8.144** 8.723Sweta x T-397 -1.488 56.290 0.583 42.207 -0.324 7.530Sweta x Neelum 3.121** 58.387 -0.745 41.647 -0.174 7.403Sweta x Mukta -5.210** 47.297 0.114 42.577 0.816 8.107Sweta x Hira -10.070** 43.567 0.445 42.227 -0.910 6.583Garima x Shekhar -0.761 54.800 1.345 41.790 -0.621 8.100Garima x Shubhra -0.098 49.527 -4.541** 35.843 -5.385** 6.557Garima x Kiran -4.916** 47.587 1.562 40.847 -0.871 7.683Garima x Surbhi -3.892** 50.007 -3.413** 36.867 0.420 8.890Garima x T-397 -1.801 54.843 0.645 40.870 -0.872 7.873Garima x Neelam 0.711 54.843 -0.200 40.793 -0.152 8.317Garima x Mukta 9.607** 60.980 -0.697 40.367 -0.706 7.477Garima x Hira -3.460* 49.043 0.377 40.757 9.515** 6.870Shekhar x Shubhra 4.145** 57.680 0.316 40.740 0.087 7.670Shekhar x Kiran -0.417 51.997 1.466 40.790 0.831 9.027Shekhar x Surbhi 0.708 56.517 0.914 41.233 0.625 8.737Shekhar x T-397 1.862 60.417 -2.378** 37.887 0.270 8.657Shekhar x Neelam 3.234* 59.277 0.401 41.433 -0.623 7.487Shekhar x Mukta -0.350 52.933 0.096 41.200 6.470** 6.353Shekhar x Hira 11.550** 42.863 2.799** 37.620 -5.026** 7.000Shubhra x Kiran 3.026** 53.503 -0.036 39.227 0.364 7.780Shubhra x Surbhi 13.091** 64.963 2.245** 42.503 1.047 8.380Shubhra x T-397 13.365** 41.253 1.153 41.357 1.171 6.437Shubhra x Neelam 6.320** 45.787 1.129 42.100 -1.077** 6.253
Table 3: Cont.....
AJEET KUMAR et al.,
213
(8.144), Surbhi×T-397 (7.127), Shekhar×Mukta (6.470),Sweta×Kiran (6.280), Sweta×Shekhar (5.847) andShubhra×Mukta (1.596) exhibited significant and positivespecific combining ability effects with good per se performancefor seed yield per plant. It clearly suggests that while selectingbest cross combinations this evidence needs to be given dueconsideration.
For instance the crosses Sweta×Shekhar had high anddesirable specific combining ability effect for days to maturity,number of capsules per plant, harvest index and seed yieldper plant; Sweta×Kiran for number of primary branches perplant, number of capsules per plant, harvest index and seedyield per plant; Sweta×Surbhi fornumber of capsules perplant, biological yield per plant and seed yield per plant;Garima×Hira for number of primary branches per plant,number of capsules per plant and seed yield per plant;Shekhar×Mukta forplant height, number of secondarybranches per plant, biological yield per plant and seed yieldper plant; Shubhra×Mukta for plant height, number of primarybranches per plant, number of capsules per plant,number ofseeds per capsule, biological yield per plant and seed yieldper plant; Shubhra×Hira for days to flowering, plant height,number of secondary branches per plant and seed yield perplant; Kiran×T-397 for number of secondary branches perplant and seed yield per plant; Surbhi×T-397 for days tomaturity, number of secondary branches per plant, biologicalyield per plant, harvest index and seed yield per plant; T-397×Neelam for harvest index and seed yield per plant; T-397×Hira for number of capsules per plant harvest indexand seed yield per plant; Neelam×Hira for days to flowering,plant height, biological yield per plant and seed yield perplant and Mukta×Hira for days to flowering, number ofprimary branches per plant, 1000 seed weight, harvest indexand seed yield per plant. These findings were supported byKalia (1972), Kaushal et al. (1974), Patil and Chopde (1983),Thakur et al. (1988), Singh et al. (1990), Khorgade et al. (1993),Popescu et al. (1995), Tiwari (1999), Goral et al.(2008), Yadav
Characters Harvest index (%) Oil content (%) Seed yield per plant (g)Crosses SCA Effect Mean SCA Effect Mean SCA Effect Mean
Shubhra x Mukta 0.413 49.760 -0.066 40.977 1.596** 8.640Shubhra x Hira -1.790 48.687 -0.318 40.040 10.967** 6.280Kiran x Surbhi -9.151** 43.600 -0.885 38.273 -0.191 7.753Kiran x T-397 -4.457** 51.040 -1.381 37.723 11.180** 7.040Kiran x Neelam 6.392** 59.377 -2.429** 37.443 -0.149 7.793Kiran x Mukta 4.191** 54.417 -1.609 38.333 -0.850 6.807Kiran x Hira 8.565** 59.920 -1.365 37.893 0.457 8.317Surbhi x T-397 4.755** 61.647 -1.413 38.687 7.127** 9.263Surbhi x Neelam 2.767* 57.147 -0.274 40.593 1.182 6.677Surbhi x Mukta 5.083** 56.703 -0.061 40.877 0.837 8.410Surbhi x Hira -0.887 51.863 -0.773 39.480 -0.985 6.790T-397 x Neelam 2.744** 59.870 -1.386 39.427 10.296** 8.430T-397 x Mukta -1.113 53.253 0.023 40.907 -0.095 7.753T-397 x Hira 5.247** 60.743 1.161 41.360 10.613** 8.663Neelam x Mukta 10.996** 62.850 -0.048 41.603 -0.121 7.450Neelam x Hira -4.247** 48.737 0.847 41.813 10.917** 8.690Mukta x Hira 2.682** 52.907 -0.074 40.963 9.183** 8.670S.E.(Sij) 2.768 1.402 0.370SE(Sij-Sik) 4.069 2.061 0.543
Table 3: Cont....
et al. (2013) and Pali and Mehta (2014).
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