Checklist of the shallow-water marine molluscs of the Azores: 1-Pico, Faial, Flores and Corvo

ACOREANA, 1998, 8(4): 487-523

CHECKLIST OF THE SHALLOW-WATER MARINE MOLLUSCS OF

Sergio P. ~ v i l a * , Jose M.N. Azevedo*, Jo2o M. Gon~alves**, Jorge Fontes*" & Frederico Cardigos*"

*SecqBo de Biologia Marinha, Departamento de Biologia, Universidade dos Aqores PT 9500 Ponta Delgada, Aqores, Portugal email: [email protected]; [email protected]

**Departamento de Oceanografia e Pescas, Universidade dos Aqores PT-9900 Horta, Aqores, Portugal

email: [email protected]; [email protected]; [email protected]

ABSTRACT An up-to-date commented check-list of the shallow-water marine molluscs of

Pico, Faial, Flores and Corvo islands (Azores, Portugal) is provided, based on literature records, on material deposited at the Department of Biology of the University of the Azores (DBUA) as a result of the scientific expeditions "Flores/89", "Aqores189" and "Pico/91", and on photographic material at the Department of Oceanography and Fisheriesluniversity of the Azores (DOPNA). The collections of

. the Museu Carlos Machado - Histdria Natural, MCM-HN, of the Museu de Zoologia - Museu de Hist6ria Natural da Faculdade de Cisncias e Tecnologia da Universidade de Coimbra, MZMHN-FCTUC, and the mollusc collection deposited at DOPIUA were surveyed and the results are included in this report.

Two hundred and fifty five taxa of shallow-water molluscs are reported to Pico, Faial, Flores and Corvo Islands (196 Gastropoda, 50 Bivalvia and 9 Cephalopoda). Of these, 32 are dubious records (23 Gastropoda, 8 Bivalvia and 1 Cephalopoda), there remaining 197 confirmed species to these islands. A total of 26 taxa were identified only down to the genus level (25 Gastropoda and 1 Bivalvia).

RESUMO Fornece-se a lista actualizada dos moluscos marinhos de iguas costeiras recolhidos

nas ilhas do Pico, Faial, Flores e Corvo, com base na bibliografia e nas amostras colhidas aquando das expediq6es cientificas "Flores/89", Aqores189" e "Pico/9lW. As fotografias submarinas de moluscos tiradas pelos bi6logos marinhos do DOPIUA - Departamento de Oceanografia e PescasIUniversidade dos Aqores (p610 da Horta, Faial), como parte do project0 "Roteiro para o turismo sub-aquitico dos Aqores" foram tambCm utilizadas neste trabalho.

As colecqBes do Museu Carlos Machado - Hist6ria Natural, MCM-HN, do Museu de Zoologia - Museu de Hist6ria Natural da Faculdade de Ciencias e Tecnologia da Universidade de Coimbra, MZIMHN-FCTUC e a colecq8o de ~noluscos existente no Departamento de Oceanografia e Pescas da Universidade dos Aqores, DOPIUA foram tambCm revistas e os resultados s5o aqui incluidos.

Duzentas e cinquenta e cinco taxa de moluscos litorais est5o dados para as ilhas do Pico: Faial, Flores e Corvo (196 Gastropoda, 50 Bivalvia e 9 Cephalopoda). Destes, 32 siio registos duvidosos (23 Gastropoda, 8 Bivalvia e 1 Cephalopoda), existindo portanto 197 espCcies confirmadas. Um total de 26 taxa foram identificados somente at6 ao gCnero (25 Gastropoda e 1 Bivalvia).

AVILA ETAL: MARINE MOLLUSCS OF THE AZORES 489

r Nordsieck & Talavera, 1979; Poppe & Goto, 1991, 1993; Macedo, 1996).

During "Aqores 89" and "Pico 91" scientific expeditions, a total of 43 and 16 samples, respectively, were collected at Pico island, from the intertidal zone to a depth of 28m. Adding the samples collected by the first author at several places on the littoral of Pico, a total of 76 samples from this island were worked. During "Aqores 89" scientific expedition, a total of 67 samples were collected at Faial island, from the intertidal zone to a depth of 30m. Recently, more samples were collected at Faial by the first author, thus making a total of 15 1 samples sorted at these two islands. During the scientific expedition "Flores 1989" and also from 27-29 October 1990, a total of 173 samples were collected around the island of Flores, from the intertidal zone to a depth of 20m (Table 1).

All molluscs were preserved in 70% alcohol and those collected during "Aqores 89" scientific expedition were sorted from algal material at the laboratories of the DOPIUA (Horta, Faial). They were later identified at the Marine Biology laboratories of the Universidade dos Aqores (Ponta Delgada, Silo Miguel). The material collected in these scientific expeditions is deposited at the Department of Biology of the University of the Azores (DBUA) under the Reference Collection numbers DBUA 178-286, DBUA 368-500, DBUA 5 17-580, DBUA 665-672, DBUA 674, DBUA 677, DBUA 678, DBUA 7 14 and DBUA

726. The samples DBUA 389, 406, 41 3,419 and 432 were lost (Table 1).

Between May 1996 and October 1997, as a part of the project "Roteiro para o Turismo Sub-Aquiitico dos Aqores", several photos of marine molluscs, especially of Opisthobranchia, were taken by the Marine Biology Team of the DOPIUA and have contributed to increase this check-list. All slides were digitized, a data bank was created at DOPIUA and the images stored in the image bank InzagDOP. Posteriorly, they were identified and labeled (Table 2). During a short visit to Faial in October 1998, the first author revised the marine molluscs that were stored at DOPIUA and the malacological reference collection was started. The identified species were also included here, under the item DOPIML (reference collection of the marine molluscs deposited at DOP) (Table 3).

The molluscs collection of the Museu de Zoologia - Museu de Hist6ria Natural da Faculdade de CiCincias e Tecnologia da Universidade de Coimbra, MZ/MHN-FCTUC, and of the Museu Carlos Machado (Hist6ria Natural), located at Ponta Delgada, MCM-HN, were surveyed and the results were also included in this work.

Species determination and the ordination of families follows the Check List of European Marine Mollusca (CLEMAM) database. The synonymy of the old citations was updated with the help of this database. Specialized works were used for the Patellidae (Titselaar,

--

1 998), Littorinidae (B ac]<e]j au Patella rnoreleti DroQt, 1858. Over rocks on the littoral of Faial (Drouet, 1858: 42).

(Nordsieck, 1977), Rissoidae (Gofas, 1930: 60) (misidentification).

& WarmOes' 1992)7 Triphoridae Patella caerulea (Linnaeus, 1758). Cais (Graham, 1988, with do Pico (Nobre, 1924: 83; 1930: 60)

extracted from CLEMAM), Turridae (misidentification). Flores (Nobre, 1924: 83;

1990; Moolenbeek & Hoenselaar, Patella vulgata Linnaeus, 1758. Laginha,

Santo Amaro, Pico (Morton et al., 1998: 61).

RESULTS Santa Cruz harbor, Flores (Morton et al., 1998: 119).

Remarks: the synonymy of the Patella of A total of 255 species of shallow- the Azores is quite complex. For further

-water are reported to the details, see Christiaens (1973), Hawkins et al. (1990) and especially Titselaar islands of Pico, Faial, Flores and (1998).

Corvo (196 Gastropods,-50 Bivalvia and 9 Cephalopoda). Of these, 31 are Patella u l~ss i~onens i s Gmelin, 1791

Patella lowei dlOrbigny, 1839. Littoral of dubious records (22 Gastropoda, 8 PiCO (Droua, 1858: 40), Bivalvia and C e ~ h a l O ~ O d a ) there Patella specrabilis Dunker, 1853. Shores are 198 confirmed species to these of Pico (Drouet, 1858: 40). islands. A total of 26 taxa were Patella i~igro-squanzosa Dunker, 1853. identified only down to the genus level Nearby Horta (Dunker, 1853: 41 fide Drouet,

(25 Gastropoda and 1 Bivalvia). 1858: 41) (misidentification). Patella baudonii Drouet, 1858. On the

rocks, near the surface of the water, at Pico (Drouet, 1858: 41).

TAXONOMIC LIST Patella caerulea var. aspera Lamarck, 18 19. Horta, over rocks of the littoral (Stn. 104, "Hirondelle", 1887) (Dautzenberg,

Phylum MOLLUSCA 1889: 67). Class GASTROPODA Patella aspera Lamarck, 18 19. Pico and

Subclass PROSOBRANCHIA Order ARCHAEOGASTROPODA

Horta, Faial and Flores (Nobre, 1924: 83; 1930: 60). Laginha, Faial (upper part of the

Family Patellidae infralittoral) (Armda & Gordo, 1984: 27).

Patella goinesi Drouet, 1858 Santa Cruz, Flores (mediolittoral) (Azevedo

Patella vulgata (Linnaeus, 1766). Azores, & Gofas, 1990: 83). at the shore (Mac Andrew, 1856: 11 8. 146) Patella caerulea (Linnaeus, 1758). Hoaa (misidentification). Pico (Nobre, 1924: 83; (Nobre, 1924: 83; 1930: 60) (misidentification). 1930: 60) (misidentification). Patella ulyssipoizensis aspera Roding,

Patella crenata Gmelin, 1790. Pico (Drouet, 1798. Reserva do Monte da Guia, Faial 1858: 40) (misidentification). (Gongalves & Martins, 1993: 32). Santo

Patella gonzesii Drouet, 1858. On the Amaro, Pico (Morton et al., 1998: 61). rocks, near the surface of the water at Pico I17zagDoP s 122-17. (Drouet, 1858: 39). DBUA 234, 667, 668. - Patella ulyssiponensis Gmelin, 179 1.

AVILA ET AL: MARINE MOLLUSCS OF THE AZORES

DBUA 179,190,240,264,265,271,458,496, dubious record because, from hundreds of 499, 554, 562, 579, 669, 670, 672, 677, 714. living specimens and shells that we have

seen from almost all the islands, none Family Acmaeidae seemed to us like Haliotis coccinea.

Acmaea virginea (O.F. Muller, 1776) ~~~t~~~ ,,irginea Miiller, 1776. A~~~~~ Haliotis tuberculata tuberculata Linnaeus,

(Jeffreys, 1882: 671). ~~~~i~ ,,irginea ~ u l l ~ ~ , 1776. B~~~~~~ Haliotis tuberculata Linnaeus, 1758. Azores,

Areia Lags and Madalena (Drou&t, 1858: 43). at shore (Mac Andrew, 1856: 119). Shores of Acnzaea virginea (Miiller, 1776). Horta Pico and Faial (Drouet, 1858: 37). Pico and

(-15 to -20 m, Stn. 103, "Hirondelle", 1887) Horta, Faial (Nobre, 1924: 82; 1930: 59-60). (Dautzenberg, 1889: 67). Pico (Nobre, 1924: Reserva do da Guia (GOn~alves 82; 1930: 60). Santa CNZ (Azevedo & Martins, 1993: 32). Faial (Wirtz, 1995: 137). Gofas, 1990: 83). Azores (Morton et al., Santa Cruz, Flores (Morton et al., 1998: 78).

H a l i o t i s t u b e r c u l a t a t u b e r c u l a t a Acnzaea virginea (O.F. Miiller, 1776). Linnaeus, 1758. IrnagDOP ~140-16,

DBUA 182, 197, 240, 274, 278, 398,410, s240-09. DOPIML 0003. DBUA 193, 197, 420, 433, 434, 457, 558, 560, 561, 562, 570, 233, 240, 263, 392, 433, 460,496,498,499, 666, 667, 668, 670, 672, 677, 726. 565, 566, 662, 668, 670, 672, 677, 726.

Family Scissurellidae Family Trochidae Anatoma crispata Fleming, 1828 Clanculus bertheloti (d70rbigny, 1840)

Scissurella crispata Fleming, 1828. Clanculus bertlzeloti (d ' o rb ign~ , 1834). Azores (Jeffreys, 1883: 88). Azores, at shore (Mac Andrew, 1856: 121, 147).

Rentarks: there are recent of this Remarks: this record needs to be confirmed. species to SHo Miguel island. Jujubinus exasperatus (Pennant, 1777) Sinezona ciizgulata (0.G.. Costa, 1861) Calliostorna (Jujubinus) exasperatus Sclzisinope fayalensis Dautzenberg, 1889. (Linnaeus, 1766). Azores (Mac Andrew,

Horta, Faial (-15 to -20 m, Stn. 103, 1856: 147). "Hirondelle", 1887) (Dautzenberg, 1889: Trocl~us eXLZSperatUS Pennant, 1777. Azores 63). Santa Cruz, Fl?res (Azevedo & Gofas, (Jeffreys, 1883: 105). 1990: 83). Pico (Avila & Azevedo, 1996: Troclzus (Ziziplzinus) exasperatus (Pennant, 106). Azores (Morton et al., 1998: 88, 89). 1777). Off Faial (-80 to -150 m, Stn. 75,

Siizezona cingulata (0 .G. Costa, 1861). "Challenger", 1873) (Watson, 1886: 696). DBUA 204, 268, 274, 276, 277,446, 465, Calliostoina (Jujubirzus) exasperatlls 466, 467, 468, 470, 471, 472, 475, 481, (Pennant, 1777). Horta (-15 to -20 m, Stn. 492, 493, 496, 499, 500, 545, 564, 662, 103, "Hirondelle", 1887) (Dautzenberg,

1889: 63). Calliostorna exasperatus (Pennant, 1777).

Family Haliotidae Cais do Pico (Nobre, 1924: 82; 1930: 59). Haliotis cocciizea Reeve, 1846 Horta (Nobre, 1924: 82; 1930: 59).

Haliotis coccinea Reeve; 1846. Littoral of Jujubinus exasperatus (Pennant, 1777). Pico and Faial (Drouet, 1858: 38). Faial Lajes do Pico (Azevedo, 1990: 59). DBUA (MZIMHN-FCTUC 11 09) (erroneously labeled 371,372,378, 388, 394, 395, 396, 400, 405, as Haliotis stonzatiaefornzis Reeve, 1846). 415,418, 421, 422, 424, 429, 432, 438, 486,

Huliotis cf. coccinea Reeve, 1843. Santa 555, 558, 560, 561, 565, 567, 568, 569, 579, Cmz, Flores (Azevedo & Gofas, 1990: 83). 671, 672, 726.

Renzarks: Nobre (1930) had doubts about Jujubinus exasperatus form tricolor the specimens of Haliotis coccir~ea listed (Pennant, 1777). Pico ( ~ v i l a L+ Azevedo, to the Azores. We also consider this a 1996: 106).

IrnagDOP s110-4, s225-19. DOPIML 0003, 0021. DBUA 557, 579, 668, 433, 670, 672.

Jujubinus pseudogravinae Nordsieck, Family Skeneidae

408,432, 441,462, 500, 556, 563, 574, 576, 1884). Horta (-15 to -20 m, Stn. 103, "Hiron- 660, 661, 662, 669, 672, 726. delie", 1887) (Dautzenberg, 1889: 46).

Jujubinus striatus (Linnaeus, 1758) Family Turbinidae Trochus striatus Linnaeus, 1758. Azores, Bolma rugosa (Linnaeus, 1767)

shore to -27 m (Mac Andrew, 1856: 120). Turbo rugosus Linnaeus, 1766. Azores, - Remarks: this record needs to be confirmed. 16 to -108m (Mac Andrew, 1856: 148).

Gibbula magus (Linnaeus, 1758) Remarks: although cited to the Azores, and living in depths studied by us - usually it lives between Gibhula lmgils Cinnaeus, 1758). Hona (-15 - 15 to -2sm (Poppe Goto, 1991) - we have to -20 m, Stn. 103, "Hirondelle", 1887)

(Dautzenbeg, 1889: 63). Pico ( ~ v i l a & never seen this species in any of the islands.

Azevedo, 1996: 106). Azores (Morton et al., Family Tricoliidae 1998: 88,144). DBUA 410,421,422,424,661,

Tricolin pullus azoriea Dautzenberg, 889 Phasiarzella pullus Linnaeus, 1758. Azores,

Calliostoma sp. -5 to -9 m (Mac Andrew, 1856: 121, 148). Calliosronla sp. Santa cruz (Azevedo & PIzasialzella azorica Dautzenberg, 1889.

Horta (-15 to -20 m, Stn. 103, "Hirondelle", 1887) (Dautzenberg, 1889: 62).

Calliostoma laugieri (Payraudeau, 1826) Phasianella pulla (Linnaeus, 1758). Cais Troclzus laugiei-i (Payraudeau, 1826). do Pico e Areia Larga (Nobre, 1924: 82;

Azores, -7 to -20m (Mac Andrew, 1856: 120, 147). 1930: 59). Horta (Nobre, 1924: 82; 1930: 59). Remarks: we consider this a dubious record. Faial (MZIMHN-FCTUC 1002).

According to Poppe & Goto (1991), Tricolia pullus Linnaeus, 1758. Azores Calliostonza laugieri is limited to the (Morton et al., 1998: 88, 89, 144). Mediterranean sea, although Nobre (1936) Tricolia pullus azorica Dautzenberg, quotes this species to Portugal, from 1889. Santa Cruz and Faj5 Grande, Flores Setubal bay south to Algarve. More (Azevedo & Gofas, 1990: 84). Lajes do Pico recently, Rolan (1984) quoted this species (Azevedo, 1990: 59). DBUA 188, 190, 193, to Ria de Vigo and Trigo Trigo & Otero 195, 197, 208, 240, 266, 270, 274,278,281, Schmitt (1987) recorded it also from Ria 286, 368, 374, 376, 377, 378,380,381, 382, de Pontevedra, Galiza (nol-th-western Spain). 384, 387, 388, 391, 393,394,395, 396, 398,

400, 403, 405, 408, 409, 410, 428, 429, 438, Calliostorna zizyphiizu~n (Linnaeus, 1758) 440, 441, 452, 453, 459, 460, 462, 465, 468,

Trochus ziz)~plzinus Linnaeus, 1758. 471, 492, 493, 496, 499, 500, 545, 551, 553, Azores ( ~ a c ~ n d r e w , 1856: 119, 147). 554, 555, 556, 557, 558, 560, 561, 563, 564,

Trochus (Zizyplzinus) zi~plzirzus Linnaeus, 566, 568, 569, 570, 571, 574, 575, 579, 662, 1758. Off Faial (-80 to -150 m, Stn. 75, 666, 667, 668, 670, 672, 677, 726. "Challenger", 1873) (Watson, 1886: 696).

Calliostorna conuloides Lamarck, 1 822. Order APOGASTROPODA Horta (-15 to -20 m, Stn. 103, "Hirondelle", Family Cerithiidae 1887) (Dautzenberg, 1889: 62). Cerithiunt zebrum Kiener, 1842

Calliostonza ~iz)~phi~zunz (Linnaeus, Ceritlziurn .zebrurn Iciener, 1842. Madale- 1758). Azores (Morton er al., 1398: 144). na, Pico (Drouet, 1858: 30).

AVILA ETAL: MARINE MOLLUSCS OF THE AZORES

Remarks: we were not able to up-date the Fossarus alnbiguus (Linnaeus, 1758). synonymy of this species. However, Madalena (Stn. 109) (Dautzenberg, 1889: Reeve (1866), gives this species to 45). Horta, Faial (Nobre, 1924: 80; 1930: Galapagos islands, so we consider this to 56). Lajes do Pico (Azevedo, 1990: 59). be a dubious record. Santa Cruz and Fajg Grande, Flores (Azevedo

& Gofas, 1990: 85). Azores (Morton et al., 1998: 89). DBUA 182, 228, 286, 387, 448,

Ceritlziunz reticulatunz (da Costa, 1778). 458, 459, 465, 471, 472, 475, 489, 492, 496, Azores, shore to - 11 5 m (Mac Andrew, 1856: 500, 661, 662, 665, 666, 677. 124, 150) (misidentification). Azores (Jeffreys, 1885: 56-57) (misidentification). Family Litiopidae

Bittium reticulatu~n (da Costa, 1778). Litiopa melanostoma Rang, 1829 Horta, Faial (-15 to -20 m, Stn. 103, Litiopa gratelupeana Drougt, 1858. "Hirondelle", 1887) (Dautzenberg, 1889: A~~~~ the ~ s a r g a s s o ~ , between the islands 41). Madalena, Areia Larga e Cais do Pica of Flares and Corvo 1858: 28-29). (Nobre, 1924: 79; 1930: 55). Horta, Faial Litiopa melanostoma Rang, 1829. Azores (Nobre, 1924: 79; 1930: 55). Lajes do Pico ( M ~ ~ ~ ~ ~ et 1998: 133). (Azevedo, 1990: 59). Azores (Morton et al., 1998: 88, 89) (misidentification). Family Triphoridae

Bittium cf. reticulatunz (da Costa, 1778). Cheirodontapallescens (Jeffreys, 1867) Santa (AzevedO Gofa% 1990: 85) Cheirodonta pallescens (Jeffreys, 1867). (misidentification). Santa Cruz, Flores (Azevedo & Gofas, 1990:

Bittium sp. DOPIML 001 0. DBUA 18 1, 182, 85). DBUA 240. 188, 190, 193,196, 197,206,240,266,274,276, 278,281,286,368,369,370,371,372,373,374, Marshallora adversa (Montagu, 1803) 375,376,377,378,379,380,381,382,384,385, Triforis pewersa (Montagu, 1803). Azores 386,387,390,391,393,394,395,396,397,398, (Mac Andrew, 1856: 150). 399,400,403,405,407,408,409,410,412,414, Triforispewersa B.D.D., 1884. Pico 415,416,417,418,422,424,427,428,430,434, (Nobre, 1930: 55). 435,436,437,438,439,441,442,443,445,446, Triphora adversa (Montagu, 1803). Santa 447,448,449,450,452,453,456,457,458,459, CNZ, Flores (Azevedo & Gofas, 1990: 85). 460,462,463,465,466,467,468,469,470,471, Lajes do Pico (Azevedo, 1990: 59). 472,474,475,476,478,480,481,483,486,489, Marshallora adversa (Montagu, 1803). 492,493,496,499,500,545,546,551,553,555, DBUA 186, 193, 197, 240, 281, 373, 374, 556,557,558,560,561,563,564,565,567,568, 379, 380, 407, 428, 441, 493, 560, 563, 574, 569,570,571,572,574,575,576,579,659,661, 662,665,666,672. 662,665, 666, 668,670,672,677,726.

Remarks: The protoconch is similar to that Monophorus erythrosolna (Bouchet & Guillemot, 1978) of Bittiunz iatreillii (Payraudeau. 1826) - Monophorus evthrosoma p o u c h e t & Bittium depauperatunz Watson, 1897

(species from Madeira and Canary islands) Guillemot, 1978). DBUA 421, 574.

and quite different from the protoconch of Mo~zophorus perversus (Linnaeus, 1758) Bittiunz reticulatu177 (da Costa, 1778) Triforis pewersa (Ljnnaeus, 1758). Azores (mainland species) (Diego Moreno, PerS. (Jeffreys, 1885: 57). Pico (Nobre, 1924: 79). comm.). For addjtional information about Hoda, ~ ~ i ~ l ( ~ ~ b ~ ~ , 1924: 79; 1930: 55). this species see A v i h & Azevedo (1997). Monoplzoms pet-versus (Linnaeus, 175 8).

Azores (Morton et al., 1998: 144). Family Fossaridae

Fossarus arnbiguus (Linnaeus, 1758) Monophorus thiriotae Bouchet, 1985 Fossarus adansoni Philippi, 1844. Azores, Monophorus tlziriotae Bouchet, 1985.

at the shore (Mac Andrew, 1856: 123,149). Azores (Bouchet, 1985: 24-26).

/

ACOREANA 1998, 8(4): 487-52

Metaxia abrupta (Watson, 1 880) Renzarks: in Morton et al. (1998: 133), the Metaxia abrupta (Watson, 1880). DBUA namesofthe Jantlzina species are switched.

Thus, where it is Janthina globosa (Fig. 7.1 -G) and Jantlzina janthina (Fig. 7.1-H),

Family Cerithiopsidae it should be Janthina globosa (Fig. 7.1-H) Cerithiopsis sp. and Janthina janthina (Fig. 7.1-G).

Cerithiopsis sp. Santa Cmz, Flores (Azevedo & Janthina pallida Thompson, 1840 Gofas, 1990: 85). Jantlzina globosa. Azores (Morton et al.,

Cerithiopsis barleei Jeffreys, 1867 1998: 133). cerithiopsis barleei ~ ~ f f ~ ~ ~ ~ , 1867, Janthina pallida Thompson, 1840. DBUA

DBUA 197,240,412,560,567. 666, 677.

Cerithiopsis jeffreysi Watson, 1885 Family Epitoniidae Cerithiopsis pulchella Jeffreys, 1858. E~itolzium algerianum (Weinkauff, 1866)

Santa cruz ( ~ ~ ~ ~ ~ d ~ & Gofas, 1990: 85). Scalariaalgeriana Weinkauff, 1866. Horta, Cerithiopsis jeffreysi Watson, 1885. Faial (-15 to -20 m, Stn. 103, "Hirondelle",

DBUA 274,574. 1887) (Dautzenberg, 1889: 56).

Cerithiopsis minima (Brusina, 1865) Epitoniurn clathratulum (Kanmacher, Ceritlziopsis cf. nzininza (Brusina, 1865).

Santa cruz (Azevedo & Gofas, 1990: 85). Scalariu clatlzratulaMontagu, 1803. ~zores,

Cerithiopsis nziniina (Brusina, 1865). -18 to -54 m (Mac Andrew, 1856: 123, 149).

Horta, Faial (-15 to -20 m, 103, cLHiron- Scalaria clatlzratula Adams, 1800. Horta,

dellen, 1887) (Dautzenberg, 1889: 43). Faial (-15 to -20 m, Stn. 103, "Hirondelle",

DBUA 188,213. 1887) (Dautzenberg, 1889: 56).

Cerithiopsis tubercularis (Montagu, 1803) Opalia crenata (Linnaeus, 1758)

Cerirhiunz tuberculare Montagu, 1803, Scalaria crelzata (Linnaeus, l758). Cais

Faial (Drouet, 1858: 31.). do Pico (Nobre, 1924: 81; 1930: 58).

Ceritlziopsis sp. Pico ( ~ v i l a & Azevedo, Family Eulimidae

Euli~na glabra (da Costa, 1778) Ceritlziopsis tubercularis (Montagu, 1803). Euli,7za suOulata Donovan, 804, Azores Azores (Morton et al., 1998: 144). DBUA 188, 379, 409, 660, 662, 666. (Jeffreys, 1884b: 365-366).

Renzarks: this record needs to be confirmed.

Family Janthinidae Parvioris sp. Ja~zthirza exigua Lamarck, 18 16 Panlioris sp. Santa Cruz, Flores (Azevedo &

Jar~tlziiza exigua Lamarck, 18 16. Azores Gofas, 1990: 85). (Mac Andrew, 1856: 119, 147). Pico (Drouet,

Vitreolina spp. Ianthina exigua Lamarck, 18 16. Horta, Viri-eolina sp. Santa Cmz, Flores (Azevedo

Faial (Nobre, 1924: 81; 1930: 58). & Gofas, 1990: 85). , MZfMHN-FCTUC 889, Viti-eolina sp. Pico (Avila & Azevedo, 1996:

106). DBUA 492,666. Ja~zthinajanthirza (Linnaeus, 1758) Ren7.ar-ks: two species are given for Flores

Janthina coinmunis Lamarck, 1816. Azores (Santa Cruz) by Azevedo & Gofas (1990). (Mac Andrew, 1856: 119, 147). Pico (Drouet, 1858: 27). Areia Larga, Pico (Nobre, 1924: Vitreolilza antifrlexa Monterosato, 1844 81; 1930: 58). Eulinza distorta Deshayes in Defrance,

Janthina janthina (Linnaeus, 1758). Azores 1823. Azores, -18 to -108 m (Mac Andrew, (Morton et al., 1998: 133). 1856: 125, 150).

AVILA ETAL: MARINE MOLLUSCS O F THE AZORES

Euliriza distorta Defrance, 1823. Azores shore (Mac Andrew, 1856: 123, 149). Pico (Jeffreys, 1884b: 367). (Drouet, 1858: 26). Madalena (Stn. 109)

Remarks: this record needs to be confirmed. (Dautzenberg, 1889: 44). Flores (Nobre, 1924: 79; 1930: 55).

Vitreolina incurva (B.D.D., 1883) Littorina striata (King, 1832). Madalena, Eulinza incurva B.D.D., 1883. Horta, Faial Areia Larga e Cais do Pico (Nobre, 1924: 79;

(-15 to -20 m, Stn. 103, "Hirondelle", 1887) 1930: 55). Horta (Nobre, 1924: 79; 1930: (Dautzenberg, 1889: 57). 55). Laginha, Faial (supralittoral) ( A e a &

Vitreolirza irzcuwa (B.D.D., 1883). DBUA . ~ ~ ~ d ~ , 1984: 27). Lajes do pica (Avila, 379, 407. 1996: 27).

Littorina striata King & Broderip, 1832. Family Littorinidae Santa Cruz, supralittoral (Azevedo & Gofas,

Melarhaphe neritoides (Linnaeus, 1758) 1990: 84). Santa Cruz (Backeljau et al., Littorina caerulescens (Lamarck, 1822). 1995: 160). Azores (Reid, 1996: 48). Santo

Pico (Drouet, 1858: 26). Amaro, Pico (Morton er al., 1998: 61, 89). Li t tor ina (Melarap l ze ) n e r i t o i d e s Santa Cruz harbor, Flores (Morton et al.,

(Linnaeus, 1758). Madalena (Stn. 109) 1998: 119). Poqa da Barra, Lajes do Pico (Dautzenberg, 1889: 44). (Morton et al., 1998: 157). DOPJML 0012.

Littorina rzeritoides (Linnaeus, 1758). DBUA 215, 221, 282, 438, 471, 661, 662, Madalena e Cais do Pico (Nobre, 1924: 663, 665, 667, 668. 79-80; 1930: 56). Horta (-15 to -20 m, Stn. 103, "Hirondelle", 1887) (Dautzenberg, Family Skeneopsidae 1889: 44); (Nobre, 1924: 79-80; 1930: 55). Skeneopsis planorbis (0 . Fabricius, 1780) Flores (Nobre, 1924: 79-80; 1930: 56). Skeneia planorbis Fabricius, 1780. Cais Laginha, Faial (supralittoral) (Arruda & do Pico (Nobre, 1924: 80; 1930: 56). Gordo, 1984: 27). Lajes do Pico (Avila, Skeneopsis aff. planorbis (Fabricius, 1780). 1996: 27). Santa Cruz (Azevedo & Gofas, 1990: 84).

Melaraplze neritoides (Ljnnaeus, 1 758). Skeneo~sis planorbis (Fabricius, 1780).

Santa cruz, nores (supraljttoral) (Azevedo Horta (-15 to -20 m, Stn. 103, "Hirondelle",

& Gofas, 1990: 84). 1887) (Dautzenberg, 1889: 46). Feteira, Melarlzaphe r~eritoides (Linnaeus, 1758). Faial (intertidal), (Chapman, 1955: 803).

Santo Amaro, Pico (Morton et al., 1998: 61). Lajes do Pico (Avila, 1996: 27). Azores (Morton et al. 1998: 88, 89, 144). Santa Cruz harbor, Flores (Morton et al., Skeneopsis jlanorbis (O.

1998: 119). DBUA 215,235,415,430,438,444, 445,448,452,457,458,459,460,496,500,659,

1780). DBUA 188,189,194,195,196,217,

660, 661, 662, 663,665,666,667,672. 268, 269, 272, 274, 277, 280, 281, 442, 444, 446, 449, 452, 456, 460,462, 466, 467, 468,

Littorina obtusata (Linnaeus, 1758) 469, 471, 475, 477: 478, 492, 493, 496, 499,

Littorina (Neritoides) obtusata (Linnaeus, 500, 564, 661, 662, 663, 674, 726.

1758). Azores (Jeffreys, 1883: 112). Remarks: we consider this a dubious record. Family Rissoidae

Rissoa guerinii RCcluz, 1843 Littori~za saxatilis (Olivi, 1792) Ciilgula costulata Alder, 1844. Feteira,

Littorina rudis (Maton, 1797). Azores Faial (mid tide) (Chapman, 1955: 803). (Jeffreys, 1883: 112-1 13). Rernarks: dubious record. This species has

Littorirla saxatilis (Olivi, 1792). Azores not been ever since cited to the Azores, id, 1996: 327). santa cruz, ~l~~~~ although its distribution ranges from the (Azevedo & Gofas, 1990: 84). DBUA 193. British isles to the Canaries ( P ~ P P ~ & Goto,

1991). Probably Chapman rnisidentified the Littorina striata King & Broderip, 1832 specimens with Rissoa guernei Dautzenberg, Littorina striata King, 1835. Azores, at 1889.

AVILA ET AL: MARWE MOLLUSCS OF THE AZORES 497

89). DBUA 188, 193, 197, 229, 240, 274, 461, 470, 474, 475, 489,490, 496, 499, 500, 409, 410, 411,421,428, 434, 438, 441, 442, 659, 660, 661, 662, 663, 665, 666, 667, 726. 444, 445,446,448, 449, 450, 451, 452, 453, 455, 456, 457, 458, 459, 460, 461, 462, 471, Crisilla postrema (Gofas, 1990) 473,474,475,4.76,483,486,489,490, 492, Setia abjecta (Watson, 1873). Ho*a (-15 493, 496, 499, 500, 551, 553, 558, 560, 561, to -20 m, Stn. 103, "Hirondelle", 1887) 564, 565, 566, 568, 570, 571, 574, 579, 659, (Dautzenberg, 1889: 52). 661, 662, 663, 665, 666, 667. Setia picta (Jeffreys, 1867). Horta (-15

to -20 m, Stn. 103, "Hirondelle", 1887) Alvania poucheti Dautzenberg, 1889 (Dautzenberg, 1889: 53).

Alvania poucheti Dautzenberg, 1889. (?) Alvania (Crisilla) sp. Santa Cruz Horta (-17 m, "Biagores", 1971) (Gofas, (Azevedo & Gofas, 1990: 85). 1990: 108). Pica (Avila &Azevedo, 1996: 106). Alvania (Crisilla) postrema Gofas, 1990. DBUA 240, 368,369, 370, 371, 372,373, 377, Horta, Faial (-3 m, -7 m, -17 m, "Biagores", 378,379,380,384,387,393,394,395,397,398, 197 1) (Gofas, 1990: 1 15). Santa Cruz, Flores 400,405,407,409,410,411,427,447,457,465, (iytertidal pool) (Gofas, 1990: 115). Pico 493,499,500,556,563,570,666. (Avila & Azevedo, 1996: 106). Azores

(Morton et al., 1998: 88, 89). Alvania sleursi (Amati, 1987) Crisilla postrerna (Gofas, 1990). DBUA

Manzonia 'leursi Amati, 1987. Bay Of 188, 198, 274, 277, 379, 384, 398,405, 412, Horta, Faial (-20 m, "Hirondelle" Stn. 193, 447, 462, 465, 470, 472, 492, 496, 499, 500, 1888) (Amati, 1987: 25-30). 545, 564, 670.

Alvania sleursi (Amati, 1987). Bay of Horta, Faial (-20 m, "Hirondelle" Stn. 193, Manzonia crassa (Kanmacher, 1798) 1888) (Gofas, 1990: 107). Santa Cruz, Flores Manzonia crassa (Icanmacher, 1798). (Gofas, 1990: 107). Pico (Avila & Azevedo, Azores (Morton et al., 1998: 88, 89). 1996: 106). DBUA 410,411,446,448,457, Remarks: we have doubts about this record. 458, 459,493, 496,499, 500, 666, 667, 726. Although some shells at first sight do

resemble Manzonia crassa, several were Cingula ordinaria Smith photographed and all showed the typical

Ciizgula ordinaria Smith. Feteira, mid-tide characters (protoconch, microsculpture of (Chapman, 1955: 803). the body whorl) of Manzonia unifasciata.

Remarks: probably this species is Ciizg~4la trifasciata ( J . Adams, 1798). ~anzonia ' unifasciata Dautzenberg, 1889

Manzonia costata (J. Adarns, 1797). Horta, Cingula trifasciata ( J . Adams, 1798) Faial (-15 to -20m, Sm. 103, "Hirondelle",

Rissoa ciizgillus (Montagu, 1803). Azores, 1887) (Dautzenberg, 1889: 5 1 ; Nobre, 1924: at shore (Mac Andrew, 1856: 122, 148). 80; 1930: 56). Pico (Nobre, 1924: 80; 1930: 56).

Cingl~la cingillus (Montagu, 1803). Pica Manzonia unifasciata Dautzenberg, 1889. (Nobre, 1924: 80; 1930: 57). Horta (Nobre, Horta, Faial (-20m, "Hirondelle", Stn. 193, 1924: 80; 1930: 57). 1888); Horta, Faial (-3m, "Bia~ores, 198 1)

Cingula trifasciata (Adams, 1798). Santa (Gofas, 1990: 116). Santa Cruz, Flores Cruz, Flores (among boulders, high in the (intertidal pools, "Flores, 1989") (Gofas, intertidal, "Flores, 1989") (Gofas, 1990: 1990: 116). Santa Cruz, Flores (Azevedo & 119). Santa Cruz, Flores (mediolittoral) Gofas, 1990: 85). Poga da Barra, Lajes do (Azevedo & Gofas, 1990: 85). Lajes do Pico Pico (Morton et al., 1998: 157). DBUA 188, (Avila, 1996: 27). FajH Grande, Flores 266, 273, 274, 281,380,381, 395, 397, 398, (Morton et al., 1998: 103). P o ~ a da Barra, 403, 409, 410,442,443, 445, 446, 449, 45 1, Lajes do Pico (Morton et al., 1998: 157). 452, 462,470,471, 475, 476, 486, 492, 493,

Cingula trifasciata (J. Adams, 1798). 496, 499, 500, 556, 571, 574, 579, 660, 661, DBUA 205,240,442,445,448,449,457,460, 662, 665, 666, 667, 670, 726.

ACOREANA 1998, 8(4): 487-52

Onoba moreleti Dautzenberg, 1889 236, 276, 468, 470, 471, 475, 492, 493, 496, Onoba moreleti Dautzenberg, 1889. Horta 499, 500, 662, 665, 666.

bay, Faial (Stn. 193) (-20 m) (Moolenbeek & Hoenselaar, 1987: 154). DBUA 181, 410, Caecum armoricum de Folin, 1869

500, 556, 666,726. Caecunz arnzoricum de Folin, 1869. Lajes do Pico (Hoeksema & Segers, 1993: 86).

Botryphallus ovummuscae (Gofas, 1990) Peringiella nitida (Brusina mss.). Horta, Caecum cf. clarkii Carpenter, 1859

~ ~ , ~ l (-15 to -20 m, stn. 103, "Hirondel]e", Caecum cf. clarkii Carpenter, 1859. Pico 1887) (Dautzenberg, 1889: 53). ( ~ v i l a & Azevedo, 1996: 106).

"Peringiella" sp. Santa Cruz, Flores (mediolittoral) (Azevedo & Gofas, 1990: 85). Caecum glabellum

Peringiella ovurnrnuscae Gofas, 1990. Caecurn Azores et ' l . ,

Santa Cruz, Flores (among boulders, high in the 1998: 89).

intertidal, " ~ l ~ ~ ~ ~ , 1989") ( ~ ~ f ~ ~ , 1990: 120). Renzarks: we could not find this species on FajZ Grande, Flores (Morton et al., 1998: 103). the CLEMAM data-base.

"Peringiella " ovunznzuscae Gofas, 1990. Pico (Avila & Azevedo, 1996: 106). Caecum vitreum Carpenter, 1859

Bofryphallus ovumnzuscae (Gofas, 1990). vitreunz Carpenter3

DBUA209,493,499,500,659.661,662,665,666. (-15 to -20 m, Stn. 103, "Hirondelle", 1887) (Dautzenberg, 1889: 43).

Family Anabathridae Pisinna glabatra (von Miihlfeldt, 1824) Family Vermetidae

Pisinna punctulurn (Philippi, 1836). Santa Vermetus cristatus Biondi, 1857

cruz, flares (intertidal pools, among algae) Verrnetus cristatus Biondi, 1859. Lajes do (Gofas, 1990: 122). Santa Cruz, Flores (Aze- Pico (Stn. 108) (Dautzenberg, 1889: 43). vedo & Gofas, 1990: 85). Lajes do Pica v~~~~~~~ triquetrus Ant. Bivona, 1832 (Avila, 1996: 27). Azores (Morton et al., 1998: 88, 89). Verrnetus sp. Santa Cruz, Flores (Azevedo &

Pisinnu glabatra (von Miihlfeldt, 1824). Gofas, 1990: 85).

DBUA 187, 189, 194, 195, 269, 272, 277, Vernzetus triquetrus Bivona-Bernardi, 1832. Pico (Avila & Azevedo, 1996: 106). 280, 442,443,444,446,447, 448,449, 450, Santa Cruz harbor, Flares et 451, 452, 456, 457, 458, 459, 460, 462, 465,

466, 467, 468, 469, 470, 471, 472, 474, 475, 1998: 119).

476, 478, 480, 481, 492, 493, 496, 499, 500, Verr7zetus triquetrus Ant. BivOna, 1832.

554, 661, 662, 663, 665, 666, 669, 726. DBUA 214,445,456,470,474,674.

Family Assimineidae Family Lamellariidae Assimi~zea eliae Paladilhe, 1875 Lamellaria latens (O.F. Miiller, 1776)

~ ~ ~ i ~ ~ i ~ ~ ~ eliae paladilhe, 1875, pico La~nellaria latens (O.F. Miiller, 1776). ( ~ v i l a & Azevedo, 1996: 106). DBUA 449, DBUA 41 0,666. 466,470, 660, 661, 665, 666, 726.

Family Triviidae Paludiitella littorina (delle Chiaje, 1828) Trivia candidula (Gaskoin, 1835) Paludinella littorina (delle Chiaje; 1828). Cypraea (Trivia) carzdidula (Gaskoin,

Santa Cruz harbor, Flores (Morton et al., 1835). Off Faial (-80 to -150 m, Stn. 75, "Challenger", 1873) (Watson, 1886: 696).

Cypraea producta Gaskoin, 1835. Family Caecidae MZIMHN-FCTUC 766.

Caecurn sp. Trivia candidula (Gaskoin, 1835). Horta, Caecunz sp. Santa Cruz, Flores (mediolittoral) Faial (Nobre, 1924: 79; 1930: 54). DBUA

(Azevedo & Gofas, 1990: 85). DBUA 212, 668, 677.

island) in the Pacific Ocean, so we consider Gordo, 1984: 27). Santa Cruz, Flores this to be a dubious record. (Azevedo & Gofas, 1990: 85). Faial (Wirtz,

1995: 143). Family Bursidae Strarnonita haenzastonza (Linnaeus,

Bursa scrobiculator (Linnaeus, 1758) 1767). Santa Cruz (Morton etal., 1998: 78). Triton scrobiculatus Lamarck, 1822. InzagDOP s123-30, s130-34, s320-22.

Azores at shore (dead) (Mac Andrew, 1856: DOP/ML 0013. DBUA 193, 233, 239, 372, 410, 422, 425, 433, 487, 576, 579, 662, 666, 668, 670, 671, 672, 677.

Between Areia Larga and Madalena (Drouet, 1858: 34). Pico (Nobre (1924: 78; 1930: 54). Family Fasciolariidae

Bursa scrobiculator (Linnaeus, 1758). Fusirzus sp. DOP/ML 0026. DBUA 495,498,672. Fusinus sp. DOPIML 28.

Family Muricidae Family Buccinidae Hexaplex trunculus (Linnaeus, 1758) Engina turbinella (Kiener, 1835)

Murex trunculus Linnaeus, 1758. Azores Engina turbinella (Kiener, 1835). Azores (Morton et al., 1998: 194). (Morton et al., 1998: 194). DBUA 410.

Hexaplex trunculus (Linnaeus, 1758). ImagDOP s155-14, s364-12, d639. DBUA425. Pisa~zia striata (Gmelin, 179 1)

Pisanianzaculosa Lamarck, 1822. Azores, Ocenebra erinacea (Linnaeus, 1758) at littoral (Mac Andrew, 1856: 133).

Murex imbricatus Brocchi. Shores of Pico ~ ~ ~ ~ ~ k ~ : this is a ~ ~ d i ~ ~ ~ ~ ~ ~ ~ ~ species (Drouet, 1858: 33). (Poppe & Goto, 1991). Dubious record.

Ocenebra erinacea (Linnaeus, 1758). DBUA 424,672. Family Nassariidae

ocinebrina aciculata ( ~ ~ ~ ~ ~ ~ k , 1822) Nassarius corniculum (Olivi, 1792) Murex corallinus Scacchi,. 1836, Azores, Nassarius corniculum (Olivi, 17g2).

-7 to -54 m (Mac Andrew, 1856: 133, 154). DBUA 418. M~~~~ aciculatus ~ ~ ~ ~ ~ ~ k , 1822. PiCO, Remarks: first recorded from SHo Miguel

dredged at -54 m (Nobre, 1924: 78; 1930: 53). (Calheta, POnta De'gada) Ocinebrina aciculata (Lamarck, 1822). Martins(l976[19801:5,13)asAm~clina

H ~ ~ ~ ~ , ~ ~ i ~ l (-15 to -20 m, stn, 103, coinicul~~rn (Olivi, 1792) this is the second ''Hirondellen, 1887) (Dautzenberg, 1889: reference to the Azores, confirming the 37). DBUA 379, 394, 395, 400, 408, 409, expansion towards west of the distribution 410,411, 672, 677, 726. of this species, formerly known to exist

only in the Mediterranean and neighboring Ocilzebrina edwardsi (Payraudeau, 1826) Atlantic waters (Algarve, Portugal) Ocinebrina edwardsi (Payraudeau, 1826). (poppe & ~ o t o , 1991).

DBUA 414. Nassarius cuvieri (Payraudeau, 1826)

Strafnolzita haemastoma (Linnaeus, 1766) Nassa costulara Brocchi, 18 14. Horta, Purpura haenzastoma (Linnaeus, 1767). Faial (Nobre, 1924: 77; 1930: 52).

Azores, at shore (Mac Andrew, 1856: 130, 153). In all the islands of the archipelago Nassarius i~zcrassatus (Strom, 1768) (Drouet, 1858: 31-32). Horta, Faial (-15 Nassa incrassata Miiller, 1776. Azores, to -20 m, Stn. 103, "Hirondelle", 1887) shore to -90 m (Mac Andrew, 1856: 13 1). (Dautzenberg, 1889: 38). Madalena, Pico Horta, Faial (-15 to -20 m, Stn. 103, (Nobre, 1924: 78; 1930: 53). "Hirondelle", 1887) (Dautzenberg, 1889:

Thais kaenzastonla (Linnaeus, 1767). 34). Very common on all the islands (Nobre, Laginha, Faial (infraiittoral) (Arruda & 1924: 77; 1930: 52). MZ-MHNIFCTUC 267.

1

1.

Volutanigra (Gmelin, 1791) = Mitra nigra 59) (misidentification). Santa Cruz, Flores (Gmelin, 1791) by Roltin, Dantart & (Azevedo & Gofas, 1990: 85) (rnisidentification). Fernandes (1997). Raphitornu carnosula (Jeffreys, 1869).

DOPML 001 1. DBUA 195, 216, 240, 274, Mitra zoizata Manyat, 18 18 278,281, 368, 372, 378, 379, 381, 387,391,

Mirra zonata Manyat, 18 18. Horta, Faial 394, 395, 397, 398, 405, 407, 408, 409, 410, h ~ b o r (-l0 m) (Bumay 19@). 428, 438, 441, 456, 496, 553, 555, 556, 561, (Avila & Azevedo, 1996: 106). Faial (Wirtz, 1995: 145). DOP/ML 0025. DBUA 495.

565, 571, 574, 579, 662, 666, 672, 677, 726.

Raphitoma linearis (Montagu, 1803) Philbertia cf. linearis (Montagu, 1803).

Santa Cruz (Azevedo & Gofas, 1990: 86). Raphitoma linearis (Montagu, 1803).

Bela septangularis Montagu, 1803. Azores (Morton et al., 1998: 144). DBUA Azores (?), shore to -14 m (Mac Andrew, 41 8, 558, 574, 666. 1856: 129, 152). DBUA 240.

Raphitoma purpurea (Montagu, 1803) Family Conidae Raphitoma purpurea (Montagu, 1803).

Azores (Morton et al., 1998: 144). DBUA 410, 666, 670.

Mitrolumna crenipicta (Dautzenberg, 1889). DBUA 418. Subclass OPISTHOBRANCHIA

Mitrolumna olivoidea (Cantraine, 1835) Order CEPHALASPIDEA Mitrolumna olivoidea (Cantraine, 1835). Family Retusidae

Lajes do Pico (Azevedo, 1990: 59). Pico Retusa multiquadrata Oberling, 1970 (Avila & Azevedo, 1996: 106). DBUA 368, Retusa mllltiquadrata Oberling, 1970. 370, 372, 375, 376, 378, 379, 380, 382, 384, Azores (Morton et al., 1998: 144). 387, 390, 393, 394, 395, 396, 397,407, 408, Ren~arks: CLEMAM cites Utriculus 410, 430, 437, 448, 455, 458, 459, 476, 486, ~ubst'iatus Jeff re~s , 1877 as the primary 488,493,496,499,500,661, 662, 677,726. reference for this taxon. However, we

follow Mikkelsen (1995: 205) to justify Bela meizkhorsti van Aartsen, 1988 the present systematic position.

Raphitonza turgidurn (Forbes, 1844). Horta, Faial (-15 to -20m, Stn. 103, "Hiron. Retusa truncatula (Bruguikre, 1792) delle", 1887) (Dautzenberg, 1889: 28). Tornatina trurtcatula (Bruguibre, 1792).

Remarks: Nordsieck (1977: 45) states that Horta, Faial (-15 to -20 m, Stn. 103, "Hiron- Bela turgida [(Forbes) Reeve, 18431 delle", 1887) (Dautzenberg, 1889: 21). (Pleurotonza) is a synonym of Pleurotorna R e t u ~ a truncatula (Bruguibre, 1792). nana Scacchi, 1836, which in turn, Azores (Morton et al., 1998: 144).

according is a 'ynonyrn of Cylichnirta umbilicata (Montagu, 1803) Bela nzenkhorsti van Aartsen, 1988. Retusa unzbilicata (Montagu, 1803). Pico Bela nebula (Montagu, 1803) (Avila & Azevedo, 1996: 106). DBUA 493,499.

Raphitoma nebula (Montagu, 1803). Dredged at -54 m, at Cais do Pico (Nobre, Family Runcinidae 1924: 76; 1930: 51). Runciiza spp.

Mallgelia nebula (Montagu, 1803). Runcina SP. Santa Cruz, Flores (~zevedo & Azores (Morton et al., 1998: 144). Gofas, 1990: 86).

Remarks: three species of Runcina are + Raphitoma carizosula (Jeffrey S, 1 869) given to Flores (Santa Cruz), by Azevedo

Anaclzis sp. Lajes do Pico (Azevedo, 1990: & Gofas (1990).


Family Bullidae Family Stiligeridae

Faial (Drouet, 1858: 24). Littoral of Pico (Wirtz, 1998: 2). (Dautzenberg, 1889: 24).

Order NOTASPIDEA Family Haminoeidae (=PLEUROBRANCOMOWHA)

Harni~zoea cf. hydatis (Linnaeus, 1758) Family Umbraculidae Hanzinoea cf. fzydatis (Linnaeus, 1758). Umbraculurn umbraculum (Roding, 1798)

DBUA 372. Umbraculunz nzediterraneum (Lamarck, 18 19). Varadouro (-2 m) (Menezes, 1991: 101).

Ha~ninoea orteai Talavera, Murillo & Urnbraculunz ur~zbraculum (Roding, Templado, 1987 1798). Seen and photographed by the first

Harninoea orteai Talavera, Murillo & author at Poqa da Baleia, south of Lajes do Templado, 1987. Azores (Morton et al., Pico, in a night dive, (14-08-1995) at -2 m.

Family Tylodinidae Family Philinidae Tylodina perversa (Gmelin, 1791)

Philine quadrata (Wood S., 1839) Tylodilza pewersa (Gmelin, 1791). Azores Philine quadrata (Wood, 1839). Off Faial (Wirtz, 1998: 2). IntugDOP s195-11, s337-30.

(-80 to -150 m, Stn. 75, "Challenger", 1873) (Watson, 1886: 696). Family Pleurobranchidae

Remarks: this species was given to Azores,

with doubts by Watson we Pleurobranchus testudinarius Cantraine, consider this a dubious record. 1840. Horta, Fajal (-15 m) (Wirtz & Martins,

1993: 56). Very common around Faial island Family Cylichnidae in a depth range of (at least) 15 to 40 m

Cylichrza alba (Brown, 1827) (Wirtz, 1995: 161; 1998: 4). Cylichna alba (Brown, 1827). Azores Pleurobranchus testudinarius Cantraine,

(Sykes, 1904: 35). 1835. OlzagDOP s128-17. DBUA 185. Remarks: we consider this a dubious record.

Berthelli~za edwardsi (Vayssibre, 1896) Cylichna cylindracea (Pennant, 1777) Berthellina sp. Santa Cruz (Azevedo &

Cyliclzna cylirzdracea (Pennant, 1777). Gofas, 1990: 86). A~~~~~ (pilsbry, 1895: 289-290; sykes, Be~tlzellina elzgeli Gardiner, 1936. Faial

(Wirtz, 1995: 163) (misidentification).

Remarks: we consider this a dubious record. Bertlzellina edwardsi (Vayssikre, 1896). Azores (up to 40m depth) (Wirtz, 1998: 4). InzagDOP s125-10, ~138-30, ~241-14.

Order SACCOGLOSSA (=ASCOGLOSSA) Order ANASPIDEA

(=APLY SIOMORPHA) Family Aplysiidae

Aplysia depilans Gmelin 1791 Apl~~sia sp. Santa Cruz, Flores (Azevedo &

Gofas, 1990: 86). Elysia orizata (Swainson, 1840) Aplysia depilans Gmelin 179 1. Faial

Elysia ornata (Swainson, 1840). 11,zagDOP (Wirtz, 1995: 157; 1998: 3). IrlzugDOP s207-26. Monte da Guia, Faial (-10 to -30 m) s139-14, s142-32, s147-30 (laying of eggs), (Wirtz, 1995: 159; 1998: 3).

Aplysia fasciata Poiret, 1798 Aplysia fasciata Poiret, 1789. Feteira (in a

rock pool) (Wirtz & Martins, 1993: 56). Clzromodoris purpurea (Laurillard, 183 1). North coast of Pico in 4 m depth (Wirtz, Faial (Wirtz, 1995: 169). Common in the 1998: 4). Seen and photographed by the first Azores, at a depth range of (at least) -4 to -20 m author at Poqa do Pano, south of Lajes do (Wirtz, 1998: 8). Pico in 18-08-95, at -1 m. ImagDOP ~139-14. CIlromodoris purpurea (Risso in GuCrin,

Aplysia parvuk Guilding in Morch, 1863 1831). Azores (Morton et al., 1998: 88,

Aplysia parvula Guilding in MGrch, 1863, 147). IlnagDoP 42, s134-149 s198-2.

Azores @ales, 1960 fide Wirtz, 1998: 16). Glossodoris edmundsi Cervera, Garcia-

Aplysia punctata (Cuvier, 1803) G6mez & Ortea, 1989

Aplysia punctata Cuvier, 1803. Faial, from G ' l o ~ ~ ~ d o ~ i s edmundsi Cervera, Garcia- 10 to 30 m depth (Wirtz, 1998: 4). Santa G6mez Ortea, lgg9. Azores (Wirtz, Cruz, Flores (Morton et al., 1998: 78). 1998: 8). InzagDOP s337-25.

Hypselodoris fonta~zdraui (Pruvot-Fol, Order NUDIBRANCHIA

Family Onchidorididae Diaphorodoris luteocincta (M. Sars, 1870)

Diaphorodoris luteocincta (Sars, 1870). da Guia (-23 m) (Wirtz, 1998:

Family Triophidae Kaloplocamus ramosus (Cantraine, 1835) 1995: 167). Azores (Wirtz, 1998: 8).

Kaloplocamus 1-antosus (Cantraine, 1835). Hypselodoris picta (Schultz in Philippi, Inside Horta harbor, Faial (-5 m) and on the 1836). I ~ ~ ~ D o ~ s126-13, ~331-34, slopes of Monte da Guia, Faial (-25 m) wrtz, 1998: 12). b7zagDOP s128-19, S139-19. Hypselodoris tricolor (Cantraine, 1835)

Hypselodoris midatlantica (Gosliner, Fanlil y Dorididae 1990). Azores (Morton et al., 1998: 88).

Doris ocelligera (Bergh, 1 88 1 ) Hypselodoris tricolor (Cantraine, 1835). Doris ocelligera (Bergh, 1881). Santa Faial (Wirtz, 1998: 9). bnagDOP s117-3.

Cruz, Flores (Azevedo & Gofas, 1990: 86). Family Discodorididae

Family Aldisidae Discodoris atrornaculata (Bergh, 1880) Aldisa biizotata Pruvot-Fol, 1953 Peltodoris atrorrlaculata Bergh, 1880.

Aldisa si7zaragdina Ortea, Perez & Llera, Azores (Morton er al., 1998: 139), 1982. Faial (down at least lam depth) Discodoris atr-onzaculata (Bergh, 1880). (Wirtz, 1998: 8).

Monte da Guia, Faial (-28 m) (Wirtz &

Family Chromodorididae Martins, 1993: 56). Faial (Wirtz, 1995: 179;

Chromodoris britoi Ortea & Perez, 1983 1998: 9). ImagDOP 39, d742. ImagDOP Cl~ronzodoris Oritoi Ortea & Perez, 1983. ~117-7, ~283-8.

Clzromodoris krohrzi (VCrany, 1846) Geitodoris cf. planata (Alder & Hancockt CIzromodoris krohni (VCrany, 1846). 1846). Santa Cruz, Flores (Azevedo & Baixa Rasa do Lajedo, Flores (InzagDOP Gofas, 1990: 86). obs). IrnagDOP 39. Discodoris planata (Alder & Hancock,

AVILA ETAL: MARINE MOLLUSCS OF THE AZORES 505

' 1846). Inside Horta harbor, Faial, in 2 and 6 m depth (Wirtz, 1998: 9).

Family Platydorididae Platydoris argo (Linnaeus, 1767)

Platydoris argo (Linnaeus, 1767). Monte da Guia, Faial (-10 to -25 m) (Wirtz & Martins, 1993: 56). Common in the Azores (Wirtz, 1998: 12). ImagDOP s139-6, s150-15.

Family Polyceridae Limaciu clavigera (O.F. Miiller, 1776) Lilnacia clavigera (Miiller, 1776). Faial

(Wirtz, 1995: 171). Azores (Wirtz, 1998: 12).

Polycera elegarzs (Bergh, 1894) Polycera elega~zs (Bergh, 1894). Horta

harbor, Faial (-5 to -25 m) (Wirtz & Martins, 1993: 56). Faial (Wirtz, 1995: 173; 1998: 12). Cabeqo do Garajau, Flores (hnagDOP obs).

Polycera quadrilineata (O.F. Miiller, 1776) Polycera quadrilineata (Miiller, 1776).

Faial, in shallow water (2 to 6m depth) (Wirtz, 1995: 171; 1998: 12). Azores (Morton et al., 1998: 144).

Family Gymnodorididae Tambja sp.

Roboastra europaea Garcia-G6mez, 1985. Faial (Wirtz, 1995: 175). IinagDOP s224-29 (misidentification).

Tanzbja sp. Pico and Faial (3 to 15 m depth) (Wirtz, 1998: 13).

Remarks: according to Wirtz (1998: 13), this is an undescribed species of the genus Tanzbja, the specimens from the Azores having been erroneously identified as Roboastra europaea Garcia-G6mez, 1985.

Tambja ceutae Garcia G6mez & Ortea, 1988

Tanzbja ceutae Garcia G6mez & Ortea, 1988. Over rocks on the shores of Faial (Wirtz & Martins, 1993: 56). Common in the Azores (from 5 to 15 m depth) (Wirtz, 1998: 12). InzagDOP s124-28, s241-28.

Family Dendrodorididae Derzdrodoris herytra ValdCz & Ortea in ValdCz, Ortea, Avila & Ballesteros, 1996 Dendrodoris grandiflora (Rapp, 1827).

Azores (Odhner, 1932) (misidentification).

Dendrodoris hei-ytra ValdCz & Ortea, 1996. Azores (ValdCs er al., 1996: 25). Outer wall of the harbor of Horta, Faial (Wirtz, 1998: 9).

Family Dotidae Doto floridicola Sirnroth, 1888

Doto Joridicola Simroth, 1888. Monte da Guia, Faial (-35 m depth) (Wirtz, 1998: 6).

Family Scyllaeidae Scyllaea pelagica Linnaeus, 1758

Scyllaea pelagica Linnaeus, 1758. Shores of the Azores, in Sargassunz (Morton et al., 1998: 136).

Family Tritoniidae Mariorziu blainvillea (Risso, 18 18)

Marionia blainvillea (Risso, 1818). Reported to the Azores, in a depth range of (at least) 4 to 15 m (Wirtz, 1995: 183; 1998: 6).

Family Facelinidae Caloria elegans (Alder & Hancock, 1845) Caloria elegans (Alder & Hancock,

1845). Monte da Guia, Faial, in a depth range of 15 to 30 m Wirtz, 1995: 183; 1998: 6).

Caloria elegans (Alder & Hancock, 1845). IllmgDOP ~135-26, ~136-26, ~266-38.

Family Fionidae Fiona pinnata (Eschscholtz, 183 1)

Fiona pirznata Alder & Hancock, 185 1. A cosmopolitan species; it is common around the Azores (Wirtz, 1998: 6). Azores (Morton et al., 1998: 133, 194).

Family Flabellinidae Flabellina pedata (Montagu, 18 15)

Flabellina pedata (Montagu, 181 5). Azores (-10 to -35m) (Wirtz, 1998: 6).

Subclass HETEROBRANCHIA Order HETEROSTROPHA

Family Rissoellidae Rissoella spp.

Rissoella sp. Santa Cruz, Flores (Azevedo & Gofas, 1990: 86).

Renlarks: two species of Rissoella are given to Flores (Santa Cruz) by Azevedo & Gofas (1 990).

Rissoella diaphana (Alder, 1848) Rissoella diaphana (Alder, 1848). Pico Graplzis sp. Santa Cruz, Flores (Azevedo &

(Avi~a & Azevedo, 1996: 106). DBUA 466, Gofas, 1990: 85).

Family Pyramidellidae Family Omalogyridae Odostomella doliolum (Philippi, 1844)

Omalogyra atomus (Philippi, 1841) Odostomiella doliolum (Philippi, 1844). Honzalogyra atornzis (Philippi, 1841). Santa Cruz, Flores (Azevedo & Gofas, 1990: 86).

Horta (-15 to -20 m, Stn. 103, "Hirondelle", Odostomella doliolum (Philippi, 1844). 1887) (Dautzenberg, 1889: 46). DBUA 574.

Ornalogyra atornus (Philippi, 1841). Lajes do Pico (~v i la , 1996: 27). Pico (Avila & Odostomia spp.

Azevedo, 1996: 106). Azores @forton er al., Brac12~stomia 'p' (Avila & Azevedo, 1996: 106).

89)' DBUA 2773 280' 446' Odostomia sp. Santa CNZ, Flores (Azevedo 449, 452, 465, 467, 468, 469, 470, 471, 474, & Gofas, 1990: 86). 475,496,499,564, 571, 662, 663,665,666, Remarks: three species of Odostomia are

given to Flores (Santa Cruz), by Azevedo Omalogyra ornata (Dautzenberg, 1889) & Gofas (1990). Hnnzalog)~ra Omata (Dautzenberg? 1889). Odostomia scalaris MacGillivray, 1843 Horta (-15 to -20 m, Stn. 103, "Hirondelle", Odostomia ,.issoides Hanley, 1844 var. 1887) (Dautzenberg, 1889: 46).

Remarks: not found on CLEMAM. alba Jeffreys, 1848. Horta, Faial (-15 to -20 m, Stn. 103, "Hirondelle", 1887)

Ammonicera sp. (Dautzenberg, 1889: 59). Ar~znzonicera sp. DBUA 199. Odostomia unidentata (Montagu, 1803) Ammoizicerafischeria~za (Monterosato, Odostonzia unidentata (Montagu, 1803).

Horta, Faial (-15 to -20 m, Stn. 103, Amrnonicera aff. jisclzeriarza (Monterosato, "Hirondelle", 1887) (Dautzenberg, 1889: 59).

1869). Santa CNZ, Flores (Azevedo & Turbonilla lactea (Linnaeus, 1758) Gofas, 1990: 84). Chenznitzia elegantissinza (Montagu,

Anzrnonicera fischeriana (Monterosato, 1803). Azores, -5 to -72 m (Mac Andrew, 1869). Pico (Avila & Azevedo, 1996: 106). 1856: 125, 150). Azores (Morton et al., 1998: 88, 89). Odostomia lactea (Linnaeus, 1758).

Azores (0 to -91 m) (Jeffreys, 1884: 357). Ammorzicera rota Hanle~ , 850) Turban illa lactea (Linnaeus, 1758). pico

Arnrnoizicera rpta (Forbes & Hanley, 1850). Pico (Avila & Azevedo, 1996: 106). (~vi la B Azevedo, 1996: 106). DBUA 493,499.

DBUA 466,470. Renzarks: although some authors (e.g.: Family Ebalidae

Fretter & Graham, 1978; Poppe & Goto, Ebala rzitidissima (Montagu, 1803) 199 1) believe that A~lznzonicer~ fischerialza Anisoc~cla nitidissima (Montagu, l803). and A. rota are the same species, Horta, Faial (-15 to -20 m, Stn. 103, "Hiron- CLEMAM lists them as different species. delle", 1887) (Dautzenberg, 1889: 56).

Family Cimidae Subclass GYMNOMORPHA Order ONCHIDIIDA

Ciina sp. Santa CNZ, Flores (Azevedo B Family Onchidiidae Gofas, 1990: 85). Onchidella celtica (Cuvier, 18 17)

Renzarks:Azevedo & Gofas (1 990) give two Onchidella,celtica (Cuvier, 18 17); Lajes species of Cinza to Flores (Santa Cruz). do Pico (Avila, 1996: 27). Pico (Avila &


Azevedo, 1996: 106). Poga da Barra, Lajes Lajinha, Faial (Nobre, 1924: 75). Horta, Faial do Pico (Morton et al., 1998: 157). DBUA (Nobre, 1924: 75; 1930: 97). Flores (Nobre, 490, 49 1, 659, 660, 661, 665. 1924: 75; 1930: 96-97) (rnisidentification).

Ovatella aequalis (Lowe, 1832). Lajes do

Subclass PULMONATA Pico (Backhuys, 1975: 56). Lajinha, Faial

Order BASOMMATOPHORA (Backhuys, 1975: 56) (misidentification).

Family Ellobiidae Ovatella vulcarzi (Morelet, 1860). Lajes do

Auriculirzella biderztata (Montagu, 1808) Pico ( ~ v i l a , 1996: 27). Faj5 Grande, Flores

Auricula alba (?) Jeffreys. Azores, at shore (Mo*On er 1998: lo3). Santa (Mac Andrew, 1856: 116). harbor, Flores (Morton et al., 1998: 119).

Leuconia bidentata Montagu, 1808. Poga da Barra, Lajes do Pico (Morton et al., Azores (Mac Andrew, 1856: 145). 1998: 157). Poga do CBo, Lajes do Pico

Auriculinella bidentata (Montagu, 1808). (Morton et al., 1998: 160). DBUA 490, 659, Santa Cmz, Flores (mediolittoral) (Azevedo 660, 661, 665, 666, 677, 726. & Gofas, 1990: 86). Lajes do Pico (Avila, Pedipes pedipes (Gmelin, 1 79 1) 1996: 27)' Fa'a Grande' er Pedipes afra FCrussac. In crevices, near the al., 1998: 103). Santa Cruz harbor, Flores (Morton et al., 1998: 119). P o ~ a da Barra, sea (Drouet, 1858: 25-26).

Lajes do Pico (Morton et al., 1998: 157). Pedipes afer Gmelin, 1790. Madalena,

Poga do C5o, Lajes do Pico (Morton et al., Pico (Stn. 109) (Dautzenberg, 1889: 19).

1998: 160). DBUA 659, 660, 661, 665, 666. Madalena, Pica (Nobre, 1924: 76; 1930: 97). Lajes do Pico and Areia Larga, Pico

Myosotella myosotis (Drapamaud, 1 80 1 ) (Backhuys, 1975: 55). Horta, Faial (Nobre, Auricula bicolor Morelet, 1860. Pico 1924: 76; 1930: 97). Lajinha, Faial

(Morelet, 1860: 206). (Backhuys, 1975: 55). Flores (Nobre, 1930: Auricula vespertina Morelet, 1860. Pico

(Morelet, 1860: 210). Pedipe~: pedipes (Gmelin, 1790). Lajes do Alexia bicolor (Morelet,. 1860). Madalena Pico (Avila, 1996: 27). P o ~ a da Barra, Lajes

and Areia Larga, Pico (Nobre, 1924: 75; do Pico (Morton et al., 1998: 157). Poga do 1930: 97). Horta, Faial (Nobre, 1924: 75; CHo, Lajes do Pico (Morton et al., 1998:

160). Faj5 Grande, Flores (Morton et al., Alexia gracilis (Lowe, 1835). Madalena, 1998: 103). Santa Cruz harbor, Flores

Pico(Nobre, 1924:75; 1930: 96). Horta, Faial (Morton et al., 1998: 119). DBUA 490, 660, (Nobre, 1924: 75; 1930: 96). 665, 677, 726.

Ovatelb ( M w o r e W lnvosotis (Drapamaud, Pseudomelampus exiguus (Lowe, 1801). Areia Larga and Cachorro, Pico (Backhuys, 1975: 57). Pseudomelanzp~s exiguus (Lowe, 183 1) .

~ ~ ~ ~ ~ ~ ~ l l ~ nll,osotis prapamaud, 1801). La~es do Pica (Avila, 1996: 27). Pica (Avila

Pico (Avila & Azevedo, 1996: 106). Faj5 & Azevedo, 1996: 106). Poqa da Barra, Lajes

Grande, Flores (Morton et al., 1998: 103). do Pico (Morton et al., 1998: 157). Poqa do

Poqa da Barra, Lajes do Pico (Morton et al., CHo, Lajes do Pico (Morton et al., 1998:

1998: 157). Poga do CBo, Lajes do Pico 160). Faj2 Grande, Flores (Morton et al.,

(Morton e t al., 1998: 160). DBUA 490, 500. 1998: 103). DBUA 659,661,665,677.

Ovatella vulcani (Morelet, 1860) Family Siphonariidae Auricula vulcarzi Morelet, 1860. Pico Williainia gussonii (da Costa, 1829)

(Morelet, 1860: 207). Patella gussonii da Costa, 1829. Azores, Marinula vulcar~i Morelet, 1860. Madalena, -22 to -36 m (Mac Andrew, 1856: 11 8).

Pico (Stn. 109) (Dautzenberg, 1889: 19). Williamia gussonii da Costa, 1829. Santa Alexia aequalis (Lowe, 1832). Madalena, Cruz, Flores (-20 m) (Azevedo & Gofas,

Pico (Nobre, 1924: 75; 1930: 96-97). 1990: 86). DBUA 240,714.

Class BNALVIA first author at S2o Miguel island and so far Subclass PTEROMORPHIA classified as T. senzigralzatus or G.

Order ARCOIDA semigranata, being so treated as distinct Family Arcidae species, do have external characteristics

Arca noae Linnaeus, 1758 that match with the descriptions of the two Arca noae Linnaeus, 1758. Azores taxa. Besides the consistent external

(Jeffreys, 1879: 571). differences in the shape of the shell, 2: Remarks: this record needs to be confirmed. semigranatus has branching periostracal

hairs on the shell, that do not appear in our Arca tetragona Poli, 1795 specimens of G. semigranata.

Arca izavicularis Bruguibre, 1792. Pico (Drouet, 1858: 45; misidentification?). Family Pinnidae

Area tetragona Poli, 1795. Azores Pinna rudis Linnaeus, 1758 (Jeffreys, 1879: 571; Morton et al., 1998: pinna rudis j-jnnaeus, 1758, A~~~~~ 89). Santa Cruz, Flores (Azevedo & Gofas, (Jeffreys, 1879: 565). Pico (Drouet, 1858: 1990: 86). zrnagDOP 39. Oo30. 45). Horta, Faia] (-15 to -20 m, Stn. 103, DBUA 240,279,666,672. "Hirondelle", 1887) (Dautzenberg, 1889:

77). Reserva do Monte da Guia, Faial Order MYTILOIDA (Gon~alves & Martins, 1993: 32). Santa

Family My tilidae Cruz, Flores (Azevedo & Gofas, 1990: 86). Mytilus edulis Linnaeus, 1758 IwwgDOP s121-18, s131-12, s140-25, s187-14.

DOPML 0016. DBUA 485,662,671.

arrives to the Azores, probably attached to the bottom of the ships and yachts, but it is not able to survive. Since there are no Order PTERIOIDA viable populations, we do not consider this Family Pteriidae

Piizctada radiata (Leach, 1 8 14) Pinctada radiata (Leach, 18 14). h a g -

DOPT319-1 to 12.

Santa Cruz, -20 m (Azevedo & Gofas, 1990: through the Suez Canal (Poppe & Goto, 86) (misidentification). 1991). It was found attached to a

Trichonzusculus senzigranatus (Reeve, ball-float, near Faial. 1858). Faial (-8 m) (Morton, 1995: 282). Azores (Morton et al., 1998: 89). DOPIML Pter%a hirundo (Linnaeus, 1758) 0017. DBUA. 193, 232, 274; 281, 398, 400, Avicula hirundo (Linnaeus, 1758). Azores 403, 412, 454, 555, 556, 570, 574, 666, 668, (Jeffreys, 1879: 565). 670, 672, 677. Avicula tarentina Lamarck, 18 19. Shores

Remarks: although on the CLEMAM of Faial (Drouet, 1858: 44). database Triclzonzusculus selnigrarzat~ls Pteria hirundo (Linnaeus, 1758). Monte (Reeve, 1858) is said to be a secondary da Guia, Faial (-25 to -50 m) (Wirtz & reference of Gi-egariella semigranata Martins, 1993: 56). Lajes do Pico (one valve (Reeve, 1858), specimens collected by the - found at shore). DOPIML 0024. DRUA 677.

b . "

C'" :


Family Pectinidae "Hirondelle", 1887) (Dautzenberg, 1889: Aequipecterz opercularis (Linnaeus, 1758)

Pecten opercularis (Linnaeus, 1758). Remarks: we were not able to up-date the Azores (Jeffreys, 1879: 558). name of this species.

Aequipecteiz opercularis Linnaeus, 1758. Horta harbor (Morton, 1967: 37). Azores Chlamys cf. varia (Linnaeus, 1758) (Morton et al., 1998: 144). Clzlamys cf. varia (Linnaeus, 1758). Horta

harbor (Morton, 1967: 37). Remarks: this record needs to be confirmed;

could be a misidentification of young

Nodipecten corallirzoides (d'orbigny, Family Anomiidae 1840). DOPIML 0018. DBUA 264, 420, Arzonzia ephippium Linnaeus, 1758 43 1,494, 672. Anomia ephippium Linnaeus, 1758.

Azores (Morton et al., 1998: 139). DBUA Cyclopectea cf. hoskynsi (Forbes, 1844) Cyclopecten cf. hoskyizsi (Forbes, 1844).

666, 672.

DOPIML 27. Heteranomia squamala (Linnaeus, 1758)

Crassadoma rnultistriata (Poli, 1795) Morzia aculeata (Miiiler, 1776). Santa Cruz,

Chlamys nzultistriata (Poli, 1795). Horta, Flores (Azevedo & Gofas, 1990: 86).

Faial (-15 to -20 m, Stn. 103, "Hirondelle", Renzarks: CLEMAM database considers

1887) (Dautzenberg, 1889: 74). Anonzia aculeata O.F. Miiller, 1776 a

Crassadoma ~izultistriata (Poli, 1795). dubious 'ynonyrn of Heteranomia In1agDOP 39. DBUA 7 14. squamala (Linnaeus, 1758).

Crassadoma pusio (Linnaeus, 1758) Family Limidae Pecten pusio (Linnaeus, 1758). Azores, Limaria hians (Gmelin, 179 1)

shore to -1'62 m (Mac Andrew, 1856: 112, Lima fragilis Scacchi. Azores, shore 143. Azores (Jeffreys, 1879: 556). to -54 m (Mac Andrew, 1856: 112, 143)

Pecten sinuosus (Gmelin, 1791). Pico (misidentification). (Drouet, 1858: 43). Lima hiaizs (Sowerby, 1820). Azores

Hinnitis sinuosus (Gmelin, 179 1). Very (Jeffreys, 1879: 564). common ill all islands (Nobre, 1924: 84; Mantellur71 hians Gmelin, 1791. Horta,

et al., 1998: 139). Faial (Nobre, 1924: 84; 1930: 62). Hinnites distortus (da Costa, 1778). Santa Limaria hiatzs (Gmelin, ]791), Reserva do Cruz (Azevedo & Gofas, 1990: 86). Crassadonza pusio (Linnaeus, 1758). Monte da Guia (Gon~alves & Martins, 1993:

32). Santa CNZ, Flores (Morton et al., 1998: DBUA 193, 2402 281, 403, 570, 666, 668, 78). ItlzagDOP ,39, 42, DOP/ML 0015, 670,672,677. DBUA 403, 574, 666, 668, 670, 671, 672,

106). DBUA 486. kimea loscombii (G.W. Sowerby I , 1824) Linza loscot7zbii (Sowerby, 1820). Azores

Chlamys rzodosa (Sowerby, 1842) (Jeffreys, 1879: 564). Chlanzys (Lyrupecten) nodosa (Sowerby, Linzea loscombii (G.W. Sowerby I, 1824).

1842). Horta, Faial (-1 5 to -20 m, Stn. 103, DBUA 403.

ACOREANA 1998, 8(4): 487-5

Order OSTREOIDA Delzada, Flores (mediolittoral) (Azevedo & Family Gryphaeidae Gofas, 1990: 87). Lajes do Pico (Avila,

Neopycnodonte cochlear (Poli, 1795) 1996: 27). Pico ( ~ v i l a & Azevedo, 1996: Ostrea cochlear Poli, 1795. Faial (Wirtz,

Neopycrzodonte cochlear (Poli, 1795). Family Montacutidae InzagDOP s147-27, s224-22. Molztacuta ferruginosa (Montagu, 1808)

Moiltacuta (Tellimya) ferrugiizosa Montagu,

Subclass HETERODONTA 1803. Horta, Faial (-15 to -20 m, Stn. 103,

Order VENEROIDA "Hirondelle", 1887) (Dautzenberg, 1889: 8 1).

Family Lucinidae Loripes lacteus (Linnaeus, 1758) Family Neoleptonidae

Loripes lucinalis (Lamarck, 18 18). DBUA Neo'eptolz "las 420, 666, 726.

Neolepton cancellatuin Salas & Gofas, Myrtea spinifera (Montagu, 1803) 1998. Flores (Santa Cruz, -20 m) (Salas &

Luciiza (Myrtea) spirzifera (Montagu, Gofas, 1998: 44-45). 1803). Azores (Jeffreys, 1881a: 701).

Remarks: there are recent records of this Family Carditidae species to SBo Miguel Island. Cardita calyculata (Linnaeus, 1758)

Cardita calyculata (Bruguibre). Azores, in Lucinorna borealis (Linnaeus, 1767) the littoral (Mac Andrew, 1856: 107,140).

L~*cina boi-ealis Linnaeus, 1767. Horta, Cardita sinllata Bruguibre, Between Areia Faial (-15 to -20 m, Stn. 103, "Hirondelle", 1887) (Dautzenberg, 1889: 59). Larga and Madalena (Drouet, 1858: 46).

Cai-dita calyculata (Linnaeus, 1758). Lucil'a borealis (Linnaeus' 758). Cais do Azores(Jeffreys, 188 l a: 705). Horta,Faial (- 15 Pico (Nobre, 1924: 86; 1930: 64). Horta, Faial (Nobre, 1924: 86; 1930: 64). to -20 m, Stn. 103, "Hirondelle", 1887)

Lucinoi7za borealis (Linnaeus, 1767). (Dautzenberg, 1889: 80). Horta, Faial Azores (Morton et al., 1998: 89). (Nobre, 1924: 85; 1930: 62). Feteira, Faial

(intertidal) (Chapman, 1955: 803). Reserva

Family Thyasiridae do Monte da Guia, Faial (Gon~alves &

Tlzyasira fluxuosa (Montagu, 1803) Martins, 1993: 32). Cais do Pico (Nobre, Axii7us flexuosus Montagu, 1803. Azores 1924: 85; 1930: 62). Poqa da Barra, Lajes do

(Jeffreys, 1881a: 701). Pico (Morton et al., 1998: 157). P o ~ a do Renzarks: this record needs to be confirmed. CBo, Lajes do Pico (Morton et al., 1998:

160). Santa Cruz and Faj5 Grande, Flores Family Lasaeidae (Azevedo & Gofas, 1990: 86). DOPIML

Lasaea ada~zsolti (Gmelin, 179 1) 0020. DBUA 181, 190, 192, 196, 240, 274, Lasaea adalzsoizi (Gmelin, 179 1 ) . Feteira 281, 368, 380, 398,403,409, 414,442,444,

(intertidal) (Chapman, 1955: 803). Poca da 445, 446, 448, 449, 451, 452, 453, 454, 457, Barra, Lajes do Pico (Morton er al., 1998: 458, 459, 460, 462, 466, 467, 468, 470, 471, 157). DBUA 179, 180,,181, 182, 188, 189, 472, 475, 480; 483, 493, 496, 499, 500, 564, 191, 192; 194, 195, 196, 197, 225, 228, 240, 568, 571, 574, 579, 662, 663, 666, 670, 671, 267, 269, 272, 274, 280, 281, 403, 442, 443, 672, 677, 726. 446, 447, 452, 453, 454, 457, 458, 459, 460, 462, 465, 466, 467, 469, 470, 471, 472, 475, Family Cardiidae 480,481,492,493,499, 500, 659, 660, 661, (?)Parvicardium exiguum (Gmelin, 1791) 662, 663, 665, 666, 726. Cardiurn pygnzaeunz Donovan, 1800.

Lasaea adansoizi (Montagu, 1808). Santa Azores, -4 to -18 m (Mac Andrew, 1856: 110). Cruz: Flores (-20 m); Faj2 Grande and Ponta Renlarks: we consider this a dubious record.

at -54 m) (Nobre, 1924: 85; 1930: 63). Horta Nototeredo lzorvegica (Spengler, 1792) harbor, Faial (Morton, 1967, Additions). Teredo rzowegica Spengler, 1792. Horta,

Faial (-15 to -20 m, Stn. 103, "Hirondelle", Globiverzus effossa (Philippi, 1836) ex 1887) (Dautzenberg, 1889: 85).

Bivona ms. Teredo rlowegica (Spengler, 1792). Cais Venus effossa (Bivona, 1836). Azores do Pico (Nobre, 1924: 86; 1930: 64). (Talisman Expedition) (Jeffreys, 1884a: 145; Nototeredo nowegica (Spengler, 1792). Nobre, 1936: 257). Azores (Morton et al., 1998: 133).

Globivenus effossa (Philippi, 1836) ex Bivona ms. DOPIML 0023. Family Xylophagidae

Timoclea ovata (Pennant, 1777) Xylophaga dorsalis (Turton, 18 19) Venus (Pasiphae) ovata Pennant, 1777. Xl'lophaga dorsalis (Turton, lg19). Horta,

Horta, Faial(-15 to -20 m, Sm. 103, "Hirondelle", Faial (-15 to -20 m, Stn. 103, "Hirondelle", 1887) (Dautzenberg, 1889: 82). Azores (Morton 1887) (Dautzenberg, 1889: 85). et al., 1998: 144). DBUA420, 714.

Gouldia minima (Montagu, 1803) Subclass ANOMALODESMATA

Gouldia minima (Montagu, 1803). Azores Order PHOLADOMYOIDA

(Morton et al., 1998: 144). DBUA 420, 672. Family Cuspidariidae Cardiomya costellata (Deshayes, 1835)

Callista chiorze (Linnaeus, 1758) C u s p i d a r i a c o s t e l l a t a D e s h a y e s , Cytlzerea chione (Linnaeus, 1758). Azores 1832. Horta bay, Faial (-20 m, Stn.29,

(sublittoral to -72 m) (Mac Andrew, 1856: 106,140). "Hirondelle", 1888) (Dautzenberg & Fischer, Ve~zns chiorze (Linnaeus, 1758). Azores (0 1897). to -219 m) (Jeffreys, 188 1a: 716).

Meretrix (Callista) cllione Linnaeus, 1758. Horta, Faial (littoral rocks) (Stn. 104, Class CEPHALOPODA "Hirondelle", 1887) (Dautzenberg, 1889: 82). Subclass COLEOIDEA Horta, Faial (Nobre, 1924: 85; 1930: 63). Order SEPIOIDEA

Callista clziorze (Linnaeus, 1758). Horta Family Spirulidae harbor, Faial (Morton, 1967, Additions). SpiruZa spirula (Linnaeus, 1758) Azores (Morton et al., 1998: 144). DBUA 420. Spirula peronii Lamarck, 1822. Azores

(Mac Andrew, 1856: 133). Pico, Areia Larga Tapes decussata (Linnaeus, 1758) (Nobre, 1924: 76; 1930: 50).

Tapes decussatus (Linnaeus, 1758). Horta harbor, Faial (Morton, 1967, Additions). Family Sepiidae

Venerzlpis pullastra (Montagu, 1803) Sepia officinalis Linnaeus, 1758

Tapes pulLastra (Montagu, 1803). Madalena, Sepia ofSicinalis Linnaeus. Pica (Drouet,

Pico (Nobre, 1924: 85; 1930: 63-64). 1858: 23). Renlarks: dubious record.

Order MYOIDA Family Hiatellidae Order TEUTHIDA

Hiatella arctica (Linnxeus, 1767) Family Loliginidae Hiatella arctica (Linnaeus, 1767). Azores Loligo forbesi Steenstrup, 1856

(Morton et al., 1998: 133, 194). Loligo vulgaris Lamarck. Pico (Drouet, 1858: 23) (misidentification).

Family Teredinidae Loligo forbesi Steenstrup, 1856. Faial Psiloteredo rnegotara (Hanley irz Forbes & (Porteiro & Martins, 1992: 11 9- 120). Azores

Hanley, 1848) (Guerra, 1992: 109-1 11). Flores (20 m depth, Teredo rnegotara Hanley, 1848. Azores Carlos Toste, pers. obs.). Ii7zagDOP s137-31,

(Morton et al., 1998: 133). s137-32, ~137-33, s137-34, s137-35.


Order OCTOPODA taxa, especially Bivalvia. Thanks are Family Octopodidae

Octopzis macropus Risso, 1826 also due to Joiio Paulo Consthcia

Octopus nzacropus Risso, 1826. ~ i t to ra~ of (MCM-HN) by the kind disposal of Faial (dephts lower than 10 m) (Gon~alves, the collections of the museum and to

Manuel Biscoito, at the Museu Octopus sahtii VCrany, 1836 Municipal do Funchal (MMF), by the

Octopus salutii VCrany, 1837. Horta, Faial facilities in the library. The first (one specimen caught in a tide pool at night) author is also grateful to the staff at (Gonqalves, 1991: 78). MZ/MHN-FCTUC for all the facilities

Octopus vulgaris Cuvier, 1797 both at the library and at the Museum, Octopus vulgaris Cuvier, 1797. Reserva do as to the facilities at the library Monte da Guia (Gonqalves & Martins, 1993: 29-33). Faial (Wirtz, 1995: 187, 193). of the Biology Department (Faculdade b7zagDOP 42. DBUA 672. de Ciencias da Universidade do Porto).

We thank the photographers who Family Tremoctopodidae

Tremoctopus violaceus violaceus delle have contributed to ImagDOP: Peter

Chiaje, 1830 Wirtz, Ricardo Serrdo Santos, and Trenzoctopus violaceus violaceus delle Carlos Taste- Travelling to Flares

Chiaje, 1830. Horta harbor and Porto Pim and Corvo were made under the beach (Gonqalves, 1991 : 76). project "Roteiro para o Turismo

Family Ocythoidae Sub-aquiitico dos Aqores", financed Ocythoe tuberculata Rafinesque, 18 14 by the Regional Government of the

0c)lthoe tuberculata Rafinesque, 1814. Azores. Thanks are also due to Horta harbor (Gonqalves, 1991: 76). ('gmara Municipal de Santa Cruz InzagDOP d 1092. (Flores), C2mara Municipal de Vila

Family Argonautidae Nova do Corvo (Corvo), Clube Argorzauta argo Linnaeus, 1758 Naval das Lajes das Flores, Museu

Argoizaiita aigo Linnaeus, 1758. Azores, das PIores, Restaurante ''0 Ca]deirgo" specially near Pico and Faial (Drouet, 1858: 21). Faial (Gonqalves, 1991: 76). b7mgDOPd983. and Associaqiio dos Bombeiros

Voluntiirios da Ilha do Corvo.

ACKNOWLEDGEMENTS The senior author was supported by the grant PRAXIS XXI/BIC/2788/96.

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Checklist of the shallow-water marine molluscs of the Azores: 1-Pico, Faial, Flores and Corvo

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