BITNER, M. A. & DULAI, A. (2004): Revision of Miocene brachiopods of the Hungarian Natural History...

Fragmenta Palaeontologica Hungarica 22, 2004

FRAGMENTA PALAEONTOLOGICA HUNGARICA 22, BUDAPEST, 2004

Revision of Miocene brachiopods of the Hungarian Natural History Museum, with special regard to the MEZNERICS collection

byMaria Aleksandra BITNER and Alfréd DULAI

Abstract — The Miocene (Karpatian and Badenian) brachiopods of the Hungarian Natural History Museum have been revised. The MEZNERICS collection comprises 16 species, Ancistrocrania abnormis (DEFRANCE), Aphelesia bipartita (BROCCHI), A. acuta MEZNERICS, Terebratula hoernesi SUESS, T. macrescens DREGER, T. styriaca DREGER, T. sinuosa (BROCCHI), T. sinuosa var. pedemontana LAMARCK, T. kemenczeiensis MAJER, Gryphusmiocaenicus (MICHELOTTI), Terebratulina retusa (LINNAEUS), Argyrotheca cordata (RISSO), Megathiris detruncata (GMELIN), Platidia anomioides (SCACCHI &PHILIPPI), Megerlia truncata (LINNAEUS) and M. margaritata MEZNERICS. The hitherto undescribed collection contains 7 species, Aphelesia sp., Terebratula sp., T. macrescens DREGER, Gryphus miocaenicus (MICHELOTTI), Argyrotheca cuneata (RISSO), A. cordata (RISSO), and Platidia anomioides (SCACCHI & PHILIPPI). The species A. cuneata is reported for the first time from the Miocene of Hungary.

Keywords — Brachiopoda, Miocene, Karpatian, Badenian, Central Paratethys, Hungary, revision.

BITNER, M. A. & DULAI, A.: Revision of Miocene brachiopods of the Hungarian Natural History Museum, with special regard to the MEZNERICScollection. — Fragmenta Palaeontologica Hungarica, 22: 69–82.

Introduction

Brachiopods played an important role during the Palaeozoic and partly in the Mesozoic but were margi-nalized in the marine communities after the end-Creta-ceous mass extinction. Palaeozoic formations are rare in Hungary; however, brachiopods are very common in the Mesozoic sedimentary formations. Sometimes they occur in rock-forming quantity, e. g. in the case of the Lower Jurassic Hierlatz Limestone in the Transdanubian Central Range (VÖRÖS 1997). Cenozoic rocks are dominant in the surface within the Carpathian Basin, but marine forma-tions were deposited only between the Eocene and the Middle Miocene. Palaeocene deposits are unknown on the surface in Hungary, while the marine connections disappeared after the Middle Miocene; the water of the Central Paratethys became brackish and later fresh water filled the Carpathian Basin (Lake Pannon).

Brachiopods are generally rare at the Eocene, Oligo-cene and Miocene localities and it is mirrored in the palaeontological literature. With the exception of a few papers, Hungarian Palaeogene and Neogene brachiopods were mainly mentioned in faunal lists only. Some new species were described by MATYASOVSZKY (1880) and MAJER (1915) but MATYASOVSZKY’s material requires a thorough revision. MEZNERICS (1943) summarized the Hungarian Tertiary brachiopod fauna in a small mono-graph on the basis of the larger Hungarian collections (Hungarian Natural History Museum, Geological Institute

of Hungary, Eötvös University, STREDA private collec-tion). The STREDA collection was later donated to the Hungarian Natural History Museum.

The aim of this paper is the revision of the Miocene brachiopods in the palaeontological collections of the Hungarian Natural History Museum. About half of the studied specimens are kept separately in the Museum as the “MEZNERICS brachiopod collection” while the others can be found in the Miocene collection arranged on the basis of the different localities. Some of these latter specimens were collected after the publication of the monograph by MEZNERICS (1943) but, surprisingly, most of them had already been in the collection before. Seemingly MEZNERICS (1943) used only a part of the available specimens but, excepting Argyrotheca cuneata, all species of the unpublished material can be found also in the “MEZNERICS collection”. A possible cause of this mixing may be the catastrophic fire in the Museum in 1956 when two-thirds of the palaeontological collections were destroyed. The undamaged material was re-arranged in the late fifties and in the sixties; the specimens might have got together during the recreation of the collection.

Both collections need taxonomical revision, as some of the specimens were identified incorrectly. There are also specimens, too badly preserved to be determined to the species level, and specimens, which are not brachio-pods at all but bivalves.

Localities and material

Karpatian — We studied 277 Karpatian (Middle Mio-cene) brachiopod specimens from 10 localities (Figure 1). All Karpatian brachiopods are from Hungary and the localities are situated within a relatively small area from

Budapest to Salgótarján. The studied Karpatian brachiopod fauna comprises 9 species.

1. Mátyásföld quarry (Budapest) — One unidentifiable terebratulid, collected by HARMAT & NOSZKY in 1927.

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702. Csömör — Eight specimens were found by STREDA

from the Anomia sand. 3. Fót — Twenty specimens were collected by HAR-

MAT, NOSZKY & SZALAI in 1927, by HARMAT & NOSZKYin 1938 and by STREDA from different points of the area (Nagyárokf ; top of Somlyó-hegy).

4. Mogyoród — One hundred and seven specimens are known from the locality. Most of them were found by NOSZKY & SZALAI in 1927 in a tributary valley NW of Mogyoród, and by NOSZKY in 1938 along the northern row of houses in the village and at Csíkvölgyi-források. Some specimens were collected by STREDA.

5. Vác — One specimen, found by NOSZKY in 1933 at Diós-hegy, along the road to Szendehely.

6. Püspökhatvan, Takács-hegy (southern trench) — With 119 specimens of 5 species, this is the richest and most diverse Karpatian brachiopod fauna in Hungary that was collected by HARMAT & NOSZKY, and by NOSZKY in

1932; three specimens are from the STREDA collection. 7. Acsa — One specimen is known from northwest of

Acsa, west of Stare Vinica-hegy. It was collected by NOSZ-KY in 1913 from sandy, tuffaceous layers in the level of the Karpatian Bryozoan Limestones.

8. Szügypuszta — Seven specimens are in the collec-tion from east of Szügy, 1 km from Leány-hegy. It was a donation by Prof. Aladár FÖLDVÁRI but originally Gyula NÁDHÁZY (a chemistry student) found the brachiopods.

9. Fels told, Aszkalinka-hegy (east side) — A single specimen was collected by NOSZKY in 1913.

10. Salgótarján — Twelve specimens from three sources: collection of HARMAT & NOSZKY in 1927 from an argilla-ceous layer next the mine dump of Károly shaft; István HRO-ZIENTSIK mine director’s donation in 1927 from the Karpa-tian Chlamys sandstone at the mine dump of Károly shaft; donation of Salgótarján Mine Museum in 1948 from the Chlamys sandstone at the sandpit above the former Casino.

Figure 1 — Sketch map showing the Karpatian localities (map modified after NAGYMAROSY & MÜLLER 1988). — a:maximum extent of Karpatian marine sediments — 1: Mátyásföld — 2: Csömör — 3: Fót — 4: Mogyoród — 5: Vác — 6: Püspökhatvan — 7: Acsa — 8: Szügypuszta — 9: Fels told — 10: Salgótarján.

Badenian — Three hundred and nine Badenian (Middle Miocene) specimens were studied from 17 localities. Most of the localities are in Hungary (mainly in the Cserhát and Börzsöny Mts) but there are several speci-mens from the Vienna Basin, from Transylvania and from Ukraine. Figure 2 shows the Hungarian outcrops as well as the localities in the Vienna Basin near the Aust-rian/Hungarian border. The studied Badenian brachio-pod fauna comprises 13 species.

1. Leitha (Lajta) Mts Austria — The mineralogist KRENNER collected 1 unidentifiable specimen in 1866.

2. Eisenstadt (Kismarton) Austria — Ninety-five specimens were studied from Hartal-hegy. Most of them

were collected by STREDA and by FRANZENAU in 1909. Some specimens were donated by the famous Austrian palaeontologist HÖRNES in 1847 and by KUBINYI in 1849 who was one of the founders of the Hungarian Geological Society. The well-known Hungarian mineralogist KREN-NER collected 3 specimens.

3. St. Margarethen (Szentmargitbánya), Austria — Two unidentifiable terebratulid specimens, found by STREDA.

4. Fert rákos — One unidentifiable terebratulid speci-men was collected by FRANZENAU in 1921.

5. Zebegény — Five specimens were found by STREDAand by NOSZKY in 1925.

6. Kemence — Eighteen specimens, STREDA collection.

Revision of Miocene brachiopods...


717. Hont — Two specimens were collected by STREDA

from the Szent János-árok. 8. Mátrasz l s — 18 specimens are known from this

locality. They were collected mostly by STREDA, by NOSZKY in 1927 from the lower quarry and by HARMATin 1927. Some specimens were donated by LEGÁNYI in 1934 and in 1937 from the quarry of Szamár-völgy.

9. Sámsonháza — Ten specimens were studied; some of them were found by STREDA at the southern side of Buda-hegy. Some other specimens were collected by STRAUSZ in 1927 and in 1931 from the Bryozoan sand of the Buda-hegy and at the eastern side of the Kiszagyva-patak, at Kiszagyva-szoros.

10. Márkháza — Fifty-three specimens, belonging to several species, derived from different sources: collec-tions by STREDA; by HARMAT & NOSZKY; by NOSZKYin 1927 from the southern and eastern sides of the Kakukk (Koklica)-hegy; by HROZIENTSIK in 1927 from the southern side of Kakukk-hegy; by MEZNERICS in 1950 from the tuffaceous limestone and tuffaceous sand layers of the Kakukk-hegy; donation of the Salgótarján Mine in 1949; donation of the Salgótarján Mine Museum.

11. Mátraverebély — Forty specimens were collected in the surroundings of this village by several geologists and palaeontologists (STREDA: Szent László-forrás; MEZNE-RICS in 1931: Remetelak; NOSZKY in 1927: western side of Szentkúti-völgy; NOSZKY in 1931: Meszes-tet and above the Szent László-forrás, lower Bryozoan level; HARMAT).

12. Nógrádszakál — Two specimens were collected by STREDA and by HARMAT, NOSZKY & SZALAI from the base layers of the Bertece-patak.

13. L pugiu (Lapugy) (Transylvania, Romania) — Two unidentifiable terebratulids by unknown collectors.

14. Co tei (Kostej) (Transylvania, Romania) — Forty-two specimens are known from this famous Transylvanian locality. Some of them were collected by STREDA, but most of them were purchased from Mrs. GERGOV in 1937.

15. Lopadea Vache (Oláhlapád) (Transylvania, Romania) — Nine specimens, collected by NOSZKY in 1929.

16. Buciumi (Törökfalu) (Transylvania, Romania) — 1 specimen by unknown collector.

17. Horodenka (Ukraine) — 8 specimens are known from the STREDA collection.

Figure 2 — Sketch map showing the Badenian localities from Hungary and the Vienna Basin (map modified after NAGYMAROSY & MÜLLER 1988). — a: maximum extent of Badenian marine sediments — 1: Leitha (Lajta) Mts — 2: Eisenstadt (Kismarton) — 3: St Margarethen (Szentmargitbánya) — 4: Fert rákos — 5: Zebegény — 6: Kemence — 7: Hont — 8: Mátrasz l s — 9: Sámsonháza — 10: Márkháza — 11: Mátraverebély — 12: Nógrádszakál.

Revision of the fauna

The MEZNERICS collection — Our revision of the Miocene brachiopods described by MEZNERICS (1943) and housed in the Hungarian Natural History Museum has shown that the following species are present (the original names used by MEZNERICS are given in bold; the results are summarized also in Table 1):

Crania abnormis (DEFRANCE in HOENINGHAUS,1828) — This species has been transferred into the genus Ancistrocrania by LEE & BRUNTON (1986). It is charac-terized by rough shell surface with concentric, irregular growth lines and anterior adductor muscle scars in form of oval elongated ridges, not united medially (Plate I: 1–3).

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72It was already recorded from the Miocene deposits of France (JULIEN 1940), Italy (DAVIDSON 1870; SACCO1902), Poland (BITNER 1990; POPIEL–BARCZYK & BAR-CZYK 1990) and Bulgaria (BITNER 1993), as well as from the Pliocene of Spain (PAJAUD 1977).

Crania subrostrata BOETTGER, 1901 — This species was originally described by BOETTGER (1901) from the Badenian of Co tei, Romania, but illustrated later by ZILCH(1934) and B RBULESCU & RADO (1984). It displays a great similarity to Ancistrocrania abnormis and probably the two species are conspecific. The similarity of these two species was already pointed out by BOETTGER (1901) and MEZNERICS (1943). However, the specimen kept in the studied collection under the number M.62.989. and named C. subrostrata, is a slightly broken dorsal valve of Megerliatruncata (LINNAEUS, 1767) (see Plate IV: 11–12).

Hemithiris bipartita (BROCCHI, 1814) — COOPER(1959) transferred this species into his new genus Aphe-lesia. The material kept in the Hungarian Natural History Museum is poorly preserved. The specimens are smaller and more elongate than typical A. bipartita (Plate I: 4–6, 7–9, 13–15). This species was also reported from the Mio-cene of Malta (PEDLEY 1976), however it is the most com-mon in the Pliocene of the Mediterranean region (GAE-TANI & SACCÀ 1985a, b; TADDEI RUGGIERO 1996; BIT-NER & MARTINELL 2001; BITNER & MOISSETTE 2003).

Hemithiris acuta MEZNERICS, 1943 — This species displays all the characters, such as smooth surface, high beak with conjunct, auriculate deltidial plates and broadly uniplicate anterior commissure, of the genus Aphelesia and should be transferred there. Aphelesia acuta is very similar to A. bipartita and these species might be synonymous, how-ever, the badly preserved material does not allow a final conclusion (Plate I: 10-12).

Hemithiris parvillima SACCO, 1902 — This species was originally described from the Miocene of Italy by SACCO (1902). Later, based on the illustrations, it was referred to as a possible fossil representative of micromor-phic rhynchonellid Cryptopora JEFFREYS (COOPER 1959; MUIR–WOOD 1959). Unfortunately, SACCO’s collection was destroyed during the World War II (FERRERO MOR-TARA et al. 1981) and its re-examination is not possible. The specimens kept under that name in the MEZNERICScollection do not represent the genus Cryptopora or any other rhynchonellids. The external characters as well the shell composed of three layers (Plate II: 27–28) indicate that those specimens belong to the genus Gryphus (GAE

TANI & SACCÀ 1984, 1985a; see also discussion in BITNER& CAHUZAC 2004), probably young representatives of G. miocaenicus (MICHELOTTI, 1847).

Terebratula MÜLLER, 1776 — MEZNERICS (1943) recognised 6 species or subspecies of Terebratula in her Mio-cene material: T. hoernesi SUESS, 1886 (Plate I: 16–17, 18–20), T. macrescens DREGER, 1889 (Plate II: 7–9, 10–12, 13–15, 16–18), T. styriaca DREGER, 1889 (Plate II: 1–3, 4–6), T. sinu-osa (BROCCHI, 1814), T. sinuosa var. pedemontana LAMARCK,1818 and T. kemenczeiensis MAJER, 1915 (Plate I: 21–23). A significant part of the material is, however, very poorly preserved and allows determination only to the generic level. The better-preserved part also does not allow esti-mating the validity of all those specific names. The speci-mens are often broken, damaged and/or crushed. None of them has a preserved loop, and even the cardinalia, visible in a few specimens, are not complete.

The problem of so-called large Terebratulas has already been discussed by several authors (COOPER 1983; GAETANI& SACCÀ 1985a; TADDEI RUGGIERO 1996). LEE et al. (2001a) summarised the complex nomenclatural history of the genus Terebratula and revised those species that occur in the Cenozoic of the Mediterranean region. They found that certainly conspecific specimens vary considerably in the degree of folding in any fossil assemblage and they may be rectimarginate to biplicate or sulciplicate. After having exa-mined their newly collected material and that kept in the Natural History Museum, London, they recognised three valid species of Terebratula: T. terebratula (LINNAEUS, 1758), T. ampulla (BROCCHI, 1814) and T. scillae SEGUENZA, 1871. The species T. sinuosa (BROCCHI, 1814) and T. calabraSEGUENZA, 1871 are placed in synonymy with T. terebratula.

All the species, except T. kemenczeiensis MAJER, 1915, reported by MEZNERICS (1943) showed a great similarity to each other and to the Miocene species T. terebratula from the Mediterranean. T. kemenczeiensis is small to medium-sized, however, much smaller than the other species discussed here. It is strongly biplicate, with folding starting very early in ontogeny. Externally it is very close to Maltaia maltensis,described by COOPER (1983) from the Miocene of Malta. However, based on the investigated material we are not able to estimate the validity of particular species and we decided to leave the original names, until a new, well pre-served and big enough material will be collected from the Miocene deposits of Hungary. As shown recently by LEE et al. (2001b), a re-evaluation of generic and specific bounda-ries for short-looped brachiopods is strongly needed. They

Explanation to Plate I

1–3 Ancistrocrania abnormis (DEFRANCE) — 5; outer, inner and lateral views of a dorsal valve, M.62.973, Mátrasz l s, Badenian. 4–6 Aphelesia bipartita (BROCCHI) — 3; dorsal, lateral and anterior views of a complete specimen; M.62.974, Márkháza, Badenian. 7–9 Aphelesia bipartita (BROCCHI) — 3; dorsal, lateral and anterior views of a complete specimen, M.62.976, Márkháza, Badenian. 10–12 Aphelesia acuta (MEZNERICS) — 3; dorsal, lateral and anterior views of a complete specimen, M.62.957, holotype, Csömör, Karpatian. 13–15 Aphelesia bipartita (BROCCHI) — 3; dorsal, lateral and anterior views of a complete specimen, M.62.959, Fót, Karpatian. 16–17 Terebratula hoernesi SUESS; 1 — 1; outer and inner views of a dorsal valve, M.62.953, Püspökhatvan, Karpatian. 18–20 Terebratula hoernesi SUESS — 1; dorsal, lateral and anterior views of a complete specimen, M.62.953, Püspökhatvan, Karpatian. 21–23 Terebratula kemenczeiensis MAJER — 1; dorsal, lateral and anterior views of a complete specimen, M.62.969, Kemence, Badenian.



73Plate I

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74took nine loop measurements of 45 specimens of Liothy-rella neozelanica and 43 specimens of L. uva, two clearly differentiated Recent brachiopod species, and found that the variation present in loops is very large and overlapping. Thus, some variation among fossil taxa, recognised as to be of specific or generic significance, should be re-interpreted.

Gryphus miocaenicus (MICHELOTTI, 1847) — This is a small species, displaying all the characters typical of the genus Gryphus (Plate II: 19–21, 22–24). The shell is smooth, biconvex with ventral valve more convex and rectimarginate anterior commissure. The pedicle foramen is small with only partly visible symphytium. The analysis of shell ultrastructure shows that the shell consists of three layers (Plate II: 27–28). The primary and secondary layers are relatively thin and composed of acicular crystallites and fibres, respectively. The tertiary layer is much thicker and built of large calcitic prisms. The presence of the tertiary layer is considered as a diagnostic generic character for Gryphus (MACKINNON & WILLIAMS 1974; GAETANI & SACCÀ 1985a). In the fossil record the genus is known since the Eocene, being, however, very common in the Pliocene of the Mediterranean region (GAETANI & SACCÀ1984, 1985a; BITNER & MARTINELL 2001; BITNER & MOISSETTE 2003). Its Recent representatives live in the Mediterranean Sea (LOGAN 1979; EMIG 1989; LOGANet al. 2004) and in the Atlantic Ocean at depths from 70 to 2663 m (BRUNTON & CURRY 1979; COOPER 1981).

Terebratulina caputserpentis (LINNAEUS, 1758) and T. caputserpentis var. granosa PONZI, 1876 — Those species are now regarded as synonyms of Terebratulina retusa(LINNAEUS, 1758) (Plate III: 1). BRUNTON et al. (1967, p. 176) re-examined the LINNAEUS’s collection and showed that LINNAEUS’s concept of the species caputserpentis varied in his own publications, and Anomia caputserpentis described by him does not belong to the genus Terebratulina. BRUN-TON et al. (1967) designated Anomia retusa as type species of the genus Terebratulina.

T. retusa is known from the Miocene and now is living in the Mediterranean Sea and the north-eastern North

Atlantic (BRUNTON & CURRY 1979; LOGAN 1979, 1993; CURRY 1982; LOGAN et al. 2004). Its depth range is very wide (from 3 to 1478 m), however, the species is most common between 100 and 500 m (CURRY 1982).

Argyrotheca subcordata (BOETTGER, 1901) and A.subcuneata (BOETTGER, 1901) — These two species are synonyms of Argyrotheca cordata (RISSO, 1826) (see also discussion in BITNER 1990: p. 141). A. cordata is typically subtriangular in outline and covered with low, almost im-perceptible ribs (Plate III: 2, 3–6, 7–8). The inner surface of margins of both valves is covered with tubercles. This species is very common in the Miocene of the Central Paratethys, being known from Austria (DREGER 1889), Bulgaria (BITNER 1993), Poland (BARCZYK & POPIEL-BARCZYK 1977; BITNER 1990, 2002; POPIEL–BARCZYK & BARCZYK 1990; BITNER & PISERA 2000; BITNER & KAIM2004), western Ukraine (FRIEDBERG 1921), Hungary (MATYASOVSZKY 1880) and Romania (BOETTGER 1901; ZILCH 1934; MEZNERICS 1943; B RBULESCU & RADO1984). In the fossil record A. cordata was also reported from the Miocene of France (JULIEN 1940), the Pliocene of Italy (DAVIDSON 1870) and Spain (ENCINAS & MARTINELL1992). Its Recent representatives live in the Mediterranean Sea (LOGAN 1979, 2003; LOGAN et al. 2004) and the Mau-ritanian region of the Atlantic Ocean (LOGAN 1983, 1988, 1993), having a depth range from few meters down to more than 600 m.

Megathiris decollata (CHEMNITZ, 1785) — The valid name for this species is Megathiris detruncata (GMELIN,1790) because it was the first binominal valid name given to this taxon (DALL 1920, p. 331). It is easily recognisable by its ribbed ornamentation and internal structures of the brachial valve, having two lateral septa (Plate III: 11–12, 13–14, 15). It is very common in the Tertiary throughout Europe (e.g. DAVIDSON 1870; MATYASOVSZKY 1880; DRE-GER 1889; SACCO 1902; JULIEN 1940; MEZNERICS 1943; PAJAUD 1977; CALZADA 1978; LLOMPART & CALZADA1982; B RBULESCU & RADO 1984; GAETANI & SACCÀ1985a; BITNER 1990; POPIEL–BARCZYK & BARCZYK

Explanation to Plate II

1–3 Terebratula styriaca DREGER — 2; dorsal, lateral and anterior views of a complete specimen, M.62.968, Zebegény, Badenian. 4–6 Terebratula styriaca DREGER — 1; dorsal, lateral and anterior views of a complete specimen, M.62.955, Fels told, Karpatian. 7–9 Terebratula macrescens DREGER — 1; dorsal, lateral and anterior views of a complete specimen, M.62.1000, Eisenstadt (Kismarton),

Badenian. 10–12 Terebratula macrescens DREGER — 1; dorsal, lateral and anterior views of a complete specimen, M.62.1000, Eisenstadt (Kismarton),

Badenian. 13–15 Terebratula macrescens DREGER — 1.5; dorsal, lateral and anterior views of a complete specimen, M.62.945, Püspökhatvan, Karpatian. 16–18 Terebratula macrescens DREGER — 1; dorsal, lateral and anterior views of a complete specimen, M.62.1000, Eisenstadt (Kismarton),

Badenian. 19–21 Gryphus miocaenicus (MICHELOTTI) — 4; dorsal, lateral and anterior views of a complete specimen, Püspökhatvan, M.62.941,

Karpatian. 22–24 Gryphus miocaenicus (MICHELOTTI) — 3; dorsal, lateral and anterior views of a complete specimen, M.62.941, Püspökhatvan,

Karpatian. 25–26 Gryphus miocaenicus (MICHELOTTI) — 25: 7; dorsal view of a complete young specimen, 26: 18; enlarged part of 25 to show details

of the posterior part, M.61.6521, Mogyoród, Karpatian. 27–28 Gryphus miocaenicus (MICHELOTTI) — 27: 380, 28: 390; transverse sections of an entire shell showing acicular primary layer

(top) underlain by fibrous secondary layer, passing into prismatic tertiary layer, M.62.962, Mogyoród, Karpatian.



75Plate II

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761990; ENCINAS & MARTINELL 1992; TADDEI RUGGIERO1996; KROH 2003), being one of the most common spe-cies in the Miocene of the Central Paratethys. Today M.detruncata is living in the Mediterranean Sea (LOGAN 1979, 2003; LOGAN et al. 2004), the eastern North Atlantic (BRUNTON & CURRY 1979; LOGAN 1983, 1988, 1993) as well as in the Caribbean Sea (COOPER 1977). It has a wide depth range from 5 to 896 m.

Platidia anomioides (SCACCHI & PHILIPPI, 1844)— P. anomioides can be easily distinguished in the investi-gated material by its characteristic large, circular foramen of amphithyridid type, bordered by very narrow deltidial plates (Plate IV: 3, 6, 8). This is a cosmopolitan species, reported from the Miocene to Recent (see DAVIDSON 1870; DREGER 1889; SACCO 1902; FOSTER 1974; COOPER 1977, 1981; BRUNTON & CURRY 1979; LOGAN 1979; B RBU-LESCU & RADO 1984; POPIEL–BARCZYK & BARCZYK1990; LOGAN et al. 2004).

Mühlfeldtia oblita (MICHELOTTI, 1839) — The name Megerlia truncata (LINNAEUS, 1767) has the priority over the name Mühlfeldtia oblita. M. truncata is easily distin-guishable by its transversely oval outline, fine ribbing and straight hinge line with a large pedicle opening (Plate IV: 13–15). It is very common both in the fossil record, being known from the Miocene (DAVIDSON 1870; DREGER 1889; SACCO 1902; FRIEDBERG 1921; MEZNERICS 1943; KOJUM-DGIEVA 1960; PAJAUD 1977; LLOMPART & CALZADA1982; B RBULESCU & RADO 1984; GAETANI & SACCÀ1985a; BITNER 1990; POPIEL-BARCZYK & BARCZYK 1990), and in the Recent waters of the Mediterranean Sea and the eastern North Atlantic (LOGAN 1979, 1983, 1988, 2003; LOGAN et al. 2004). It belongs to the eurybathic forms living from 60 to 600 meters, however, it also occurs occa-sionally in shallow caves where environmental conditions are similar to those in deeper water (LOGAN 1979, 2003).

Mühlfeldtia margaritata MEZNERICS, 1943 — This species should be now transferred to the genus Megerlia. M. margaritata is very close to M. truncata, differing in having more distinct ribs and shorter beak, however, this last dif-ference might be due to the state of preservation (Plate IV: 16–18). The very limited, not well-preserved material pre-vents any detailed investigations.

The presence of this species was also recorded from the

Badenian of Romania by B RBULESCU & RADO (1984). However, the specimen illustrated by B RBULESCU & RADO (1984, pl. 3, figs. 13a, 13c), although partly broken, clearly represents Megerlia truncata. The re-examination of the material from the type locality is needed to resolve this taxonomic problem.

Undescribed collection — In the Museum’s Miocene material there are also several undescribed specimens, however with some preliminary determination. Unfortu-nately, in some cases those determinations are erroneous or the material is very poorly preserved not allowing identification at the specific level (see Table 1). In four cases, the name Terebratula grandis BLUMENBACH, 1803 is used for big, smooth Terebratula. T. grandis was originally described from the Oligocene of Germany and now this species was transferred to the north European genus Plio-thyrina (ROY 1980; COOPER 1983). Pliothyrina resembles Terebratula externally but differs internally in the presence of well-developed inner hinge plates, which are absent in Terebratula. There are also specimens in the collection determined as Terebratula sinuosa. This Tertiary Mediterra-nean species is now included into the synonymy of another Mediterranean species T. terebratula (LINNAEUS) by LEE et al. (2001a). It is difficult to say if the Hungarian material belongs to T. terebratula or to any of the species described from the Central Paratethys by DREGER (1889). Like in the MEZNERICS collection, we decided to leave the original names, having no possibility to make a real revision of the poorly preserved material.

The taxonomic composition of the undescribed specimens is very similar to that published by MEZ-NERICS (1943). We recognised the following valid spe-cies:

Aphelesia sp. — Seven specimens from 2 localities (see details in Table 1).

Terebratula sp. — More than 70 specimens from 11 localities (see details in Table 1).

Terebratula macrescens DREGER, 1889 — Fifteen specimens from 3 localities (see details in Table 1).

Gryphus miocaenicus (MICHELOTTI, 1847) — More than 100 specimens from 6 localities (see details in Table 1) (Plate II: 25–26).

Explanation to Plate III

1 Terebratulina retusa (LINNAEUS) — 10; dorsal view of a complete specimen, M.62.943, Püspökhatvan, Karpatian. 2 Argyrotheca cordata (RISSO) — 20; dorsal view of a complete specimen, M.62.986, Co tei (Kostej), Badenian. 3–6 Argyrotheca cordata (RISSO) — complete specimen — 3: 20; dorsal view — 4: 30; enlarged part of 3 to show details of the

posterior part — 5: 25; lateral view — 6: 50; enlarged part of 5 to show coarsely punctate surface, M.62.988, Co tei (Kostej), Badenian. 7–8 Argyrotheca cordata (RISSO) — complete specimen — 7: 22; dorsal view — 8: 42; enlarged part of 7 to show details of the

posterior part, M.62.987, Co tei (Kostej), Badenian. 9 Argyrotheca cuneata (RISSO) — 16; dorsal view of a complete specimen, M.62.3883, Mátrasz l s, Badenian. 10 Argyrotheca cuneata (RISSO) — 20; dorsal view of a complete specimen, M.62.3883, Mátrasz l s, Badenian. 11–12 Megathiris detruncata (GMELIN) — 15; inner and lateral views of a dorsal valve, showing medium septum, two lateral septa, and

rudimentary fourth and fifth septum, M.62.984, Co tei (Kostej), Badenian. 13–14 Megathiris detruncata (GMELIN) — complete specimen — 13: 16; dorsal view — 14: 120; fragment of the surface with numerous

puncta and fibres of the secondary layer visible, M.62.984, Co tei (Kostej), Badenian. 15 Megathiris detruncata (GMELIN) — 11; dorsal view of a complete specimen, M.62.984, Co tei (Kostej), Badenian.



77Plate III

BITNER & DULAI


78Argyrotheca cuneata (RISSO, 1826) — Two speci-

mens from Mátrasz l s (Plate III: 9, 10). This species has not been recognised in the MEZNERICS collection, however it was already described from the Miocene of Hungary under the name Argiope hofmanni by MATYASOVSZKY (1880). A. cuneata is well-known from the Middle Miocene deposits of the Central Paratethys (BITNER 1990, 1993; POPIEL–BARCZYK & BARCZYK 1990; BITNER & PISERA 2000; KROH 2003; BITNER & KAIM 2004). Today it is living in the Mediterranean Sea (LOGAN 1979, 2003; LOGAN et al.

2004) and the eastern North Atlantic (BRUNTON & CURRY 1979; LOGAN 1983, 1988, 1993). Its depth range is similar to that of A. cordata, from few meters to more than 600 m (LOGAN 1979, 1983).

Argyrotheca cordata (RISSO, 1826) — Three speci-mens from one locality (see details in Table 1).

Platidia anomioides (SCACCHI & PHILIPPI, 1844)— Twenty-five specimens from 4 localities (see details in Table 1) (Plate IV: 1, 2–4, 5–6, 7–8, 9–10).

* * * Acknowledgements — This paper is a contribution to the joint Hungarian-Polish Project 27 Miocene Paratethyan brachiopod and molluscan fauna: environmental and climatic implications realised within the framework of bilateral co-operation between the Hungarian Academy of Sciences and Polish Academy of Sciences. We thank J. PÁLFY for the revision of the English text. The paper benefited from critical review by A. VÖRÖS. The SEM micrographs were taken in the SEM laboratory of the Institute of Paleobiology (Warszawa) using Philips XL-20 scanning electron microscope. D. SZABÓ took the photos of the macroscopic specimens.

References

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BARCZYK, W. & POPIEL–BARCZYK, E. (1977): Brachiopods from the Korytnica basin (Middle Miocene; Holy Cross Mountains, Poland). — Acta Geologica Polonica, 27: 157–167.

BITNER, M. A. (1990): Middle Miocene (Badenian) brachiopods from the Roztocze Hills, south-eastern Poland. — Acta Geologica Polonica,40: 129–157.

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BRUNTON, C. H. C., COCKS, L. R. M. & DANCE, S. P. (1967): Brachiopods in the Linnaean Collection. — Proceedings of the Linnean Society of London, 178: 161–183.

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Explanation to Plate IV

1 Platidia anomioides (SCACCHI & PHILIPPI) — 17; dorsal view of a complete specimen, M.62.3142, Mátraverebély, Badenian. 2–4 Platidia anomioides (SCACCHI & PHILIPPI) — a complete specimen — 2: 15; dorsal view — 3: 32; enlarged part of 2 to

show details of the posterior part — 4: 15; lateral view, M.62.3142, Mátraverebély, Badenian. 5–6 Platidia anomioides (SCACCHI & PHILIPPI) — a complete specimen — 5: 10; dorsal view — 6: 18; enlarged part of 5 to

show details of the posterior part, M.62.3051, Mátraverebély, Badenian. 7–8 Platidia anomioides (SCACCHI & PHILIPPI) — ventral valve — 7: 10; inner view — 8: 24; enlarged part of 7 to show details

of the posterior part, M.62.3051, Mátraverebély, Badenian. 9–10 Platidia anomioides (SCACCHI & PHILIPPI) — dorsal valve — 9: 12; inner view — 10: 22; enlarged part of 9, visible details

of the posterior part, M.62.3143, Mátraverebély, Badenian. 11–12 Megerlia truncata (LINNAEUS) — dorsal valve of a young specimen — 11: 14; inner view — 12: 400; enlarged part of the shell

surface showing secondary layer fibres, M.62.989, Co tei (Kostej), Badenian. 13–15 Megerlia truncata (LINNAEUS) — 4.5; dorsal, lateral and anterior views of complete specimen, M.62.938, Püspökhatvan,

Karpatian.16–18 Megerlia margaritata (MEZNERICS) — 4.5; ventral, lateral and anterior views of complete specimen, M.62.993, Buciumi

(Törökfalu), Badenian.



79Plate IV

BITNER & DULAI


80DREGER, J. (1889): Die tertiären Brachiopoden des Wiener Beckens. —

Beiträge zur Paläontologie Oesterreich–Ungarns, 7: 179–192. EMIG, C. C. (1989): Distributional patterns along the Mediterranean

continental margin (Upper Bathyal) using Gryphus vitreus (Brachiopoda) densities. — Palaeogeography, Palaeoclimatology, Palaeoecology, 71: 253–256.

ENCINAS, A. & MARTINELL, J. (1992): Braquiopódos del Pliocene del Emporda (Catalunya). — III Congreso Geologico de Espana y VIII Congreso Latinoamericano de Geologia, Salamanca 1992, 1: 501–505.

FERRERO MORTARA, E., MONTEFAMEGLIO, L., PAVIA, G. & TAMPIERI,R. (1981): Catalogo dei tipi e degli esemplari figurati della collezione BELLARDI e SACCO. Parte I. — Cataloghi Museo Regionale di Scienze Naturali, VI, 327 pp.

FOSTER, M. W. (1974): Recent Antarctic and Subantarctic brachiopods. — Antarctic Research Series, 21: 1–189.

FRIEDBERG, W. (1921): Les brachiopodes miocènes de la Podolie Occidentale. — Prace Naukowe Uniwersytetu Pozna skiego, Sekcja Matematyczno–Przyrodnicza, 2: 1–20.

GAETANI, M. & SACCÀ, D. (1984): Brachiopodi batiali nel Pliocene e Pleistocene di Sicilia e Calabria. — Rivista Italiana di Paleontologia e Stratigrafia, 90: 407–458.

GAETANI, M. & SACCÀ, D. (1985a): Brachiopodi neogenici e pleistocenici della provincia di Messina e della Calabria meridionale. — Geologica Romana, 22 (1983): 1–43.

GAETANI, M. & SACCÀ, D. (1985b): Il genere Aphelesia (Rhynchonellida, Brachiopoda) nel Mio-Pliocene italiano. — Rivista Italiana di Paleontologia e Stratigrafia, 91: 357–378.

JULIEN, M. (1940): Révision de la faune vindobonienne de Sain-Fons (Rhône). — Travaux du Laboratoire de Géologie de la Faculté des Sciences de Lyon, 38: 1–60.

KOJUMDGIEVA, E. (1960): In: KOJUMDGIEVA, E. & STRACHIMIROV, B. (Eds.): Les fossiles de Bulgarie; VII, Tortonien, Académie des Sciences de Bulgarie, Sofia, pp.1–246.

KROH, A. (2003): The Brachiopoda of the Langhian (Lower Badenian) of the Molasse Zone and the northern Vienna Basin (Austria). — Annalen des Naturhistorischen Museums in Wien, 104A: 145–153.

LEE, D. E. & BRUNTON, C. H. C. (1986): Neocrania n. gen., and a revision of Cretaceous–Recent brachiopod genera in the family Craniidae. —Bulletin of the British Museum (Natural History), Geology, 40: 141–160.

LEE, D. E., BRUNTON C. H. C., TADDEI RUGGIERO, E., CALDARA, M. & SIMONE, O. (2001a): The Cenozoic brachiopod Terebratula: its type species, neotype, and other included species. — Bulletin of the Natural History Museum, Geology Series, 57: 83–93.

LEE, D. E., CARLSON, S. J., BUENING, N. & SAMSON, C. R. (2001b):Variation in the loops of two Recent species of Liothyrella(Brachiopoda; Terebratuloidea) from New Zealand and South Orkney Islands. — In: BRUNTON, C. H. C., COCKS, L. R. M., LONG,S. L. (Eds): Brachiopods past and present. — The Systematics Association Special Volume Series, 63: 56–70.

LLOMPART, C. & CALZADA, S. (1982): Braquiópodos messiniensis de la isla de Menorca. — Boletin de la Real Sociedad Española de Historia Natural, 80: 185–206.

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LOGAN, A. (1988): Brachiopoda collected by CANCAP IV and VI expeditions to the south-east North Atlantic. 1980–1982. — Zoo-logische Mededelingen Leiden, 62: 59–74.

LOGAN, A. (1993): Recent brachiopods from the Canarian-Cape Verdean region: diversity, biogeographic affinities, bathymetric range and life habits. — Courier Forschungsinstitut Senckenberg, 159: 229–233.

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Authors’ addresses: Dr. Maria Aleksandra BITNERInstitute of Paleobiology Polish Academy of Sciences ul. Twarda 51/55 00–818 Warszawa, Poland E-mail: [email protected]

Dr. Alfréd DULAIGeological and Palaeontological Department Hungarian Natural History Museum Map: Budapest, VIII, Ludovika tér 2 Mail: 1431 Budapest, pf. 137

Hungary E-mail: [email protected]



81Appendix

Table 1 — The revised Miocene (Karpatian and Badenian) brachiopod fauna of the Hungarian Natural History Museum. — C: complete specimen — D: dorsal valve — V: ventral valve — F: fragment.

The MEZNERICS collection

Inventory Old name New name Material Locality Age

M.62.938. Mühlfeldtia oblita Megerlia truncata 3C, 4V Püspökhatvan Karpatian M.62.939. Mühlfeldtia oblita Megerlia truncata 2C, 1V, 1D Püspökhatvan Karpatian M.62.940. Gryphus miocaenicus Gryphus miocaenicus

Megerlia truncata 12C, 2V 1D

Püspökhatvan Karpatian

M.62.941. Gryphus miocaenicus Gryphus miocaenicus 18C, 8V Püspökhatvan Karpatian M.62.942. Gryphus miocaenicus Gryphus miocaenicus 2C Püspökhatvan Karpatian M.62.943. Terebratulina caputserpentis var. granulosa Terebratulina retusa 1C Püspökhatvan Karpatian M.62.944. Terebratulina caputserpentis Terebratulina retusa

Gryphus miocaenicus 5C, 2V 1C

Püspökhatvan Karpatian

M.62.945. Terebratula macrescens Terebratula macrescens 2C, 5V Püspökhatvan Karpatian M.62.946. Terebratula macrescens Terebratula macrescens 1C, 2V Püspökhatvan Karpatian M.62.947. Terebratula hoernesi Terebratula hoernesi 2C, 1V Püspökhatvan Karpatian M.62.948. Terebratula hoernesi juv. Terebratula hoernesi 1D, 1F Püspökhatvan Karpatian M.62.949. Terebratula hoernesi Terebratula hoernesi 3C, 13V, 2D Püspökhatvan Karpatian M.62.950. Terebratula hoernesi Terebratula hoernesi 2C Püspökhatvan Karpatian M.62.951. Terebratula hoernesi Terebratula hoernesi 1C, 1V, 2D Püspökhatvan Karpatian M.62.952. Terebratula hoernesi Terebratula hoernesi 1V, 1D Püspökhatvan Karpatian M.62.953. Terebratula hoernesi Terebratula hoernesi 2C, 1D Püspökhatvan Karpatian M.62.954. Terebratula hoernesi Terebratula hoernesi 1C, 2F Püspökhatvan Karpatian M.62.955. Terebratula styriaca Terebratula styriaca 1C Fels told Karpatian M.62.956. Hemithiris acuta Aphelesia acuta 2C, 2V, 2D Csömör Karpatian M.62.957. Hemithiris acuta (holotype) Aphelesia acuta 1C Csömör Karpatian M.61.4375. Hemithiris acuta Aphelesia acuta 1V Csömör Karpatian M.62.958. Terebratula sinuosa var. pedemontana Terebratula sinuosa var. pedemontana 1C Acsa Karpatian M.62.959. Terebratula sinuosa var. pedemontana Terebratula sp.

Aphelesia bipartita 1F1C

Fót Karpatian

M.62.960. Terebratula macrescens Terebratula macrescens 1V Fót Karpatian M.62.962. Hemithiris parvillima Gryphus miocaenicus 27C, 5V, 1D Mogyoród Karpatian M.62.963. Gryphus miocaenicus Gryphus miocaenicus 4C Mogyoród Karpatian M.62.964. Terebratula macrescens Terebratula macrescens 6C, 1V Szügypuszta Karpatian M.62.965. Terebratula sinuosa Terebratula sinuosa 1D Sámsonháza Badenian M.62.966. Terebratula kemenczeiensis Terebratula kemenczeiensis 5C, 1V Sámsonháza Badenian M.62.967. Gryphus miocaenicus Gryphus miocaenicus 4C Zebegény Badenian M.62.968. Terebratula styriaca Terebratula styriaca 1C Zebegény Badenian M.62.969. Terebratula kemenczeiensis Terebratula kemenczeiensis 3C Kemence Badenian M.62.970. Terebratula kemenczeiensis Terebratula kemenczeiensis 5C, 2V, 2D Kemence Badenian M.62.971. Gryphus miocaenicus Gryphus miocaenicus 2C, 1V Kemence Badenian M.62.972. Terebratula Terebratula sp. 2C, 1D Kemence Badenian M.62.961. Terebratula sinuosa juv. Terebratula sinuosa juv. 2V Hont Badenian M.62.973. Crania abnormis Ancistrocrania abnormis 3D Mátrasz l s Badenian M.62.974. Hemithyris bipartita Aphelesia bipartita 2C Márkháza Badenian M.62.975. Hemithyris bipartita Aphelesia bipartita 2C Márkháza Badenian M.62.976. Hemithyris bipartita Aphelesia bipartita 2C Márkháza Badenian M.62.977. Terebratula sinuosa Terebratula sinuosa 1C Márkháza Badenian M.62.978. Terebratula macrescens Terebratula macrescens 1C Márkháza Badenian M.62.979. Platidia anomioides Platidia anomioides 5C, 4V, 1D Mátraverebély Badenian M.62.980. Terebratula macrescens Terebratula macrescens 2C, 5V Mátraverebély Badenian M.62.981. Terebratula cf. macrescens Terebratula cf. macrescens 1V Mátraverebély Badenian M.62.982. Terebratula sinuosa Terebratula sinuosa 1V Nógrádszakál Badenian M.62.983. Terebratula sp. Terebratula sp. 2D L pugiu Badenian M.62.984. Megathyris decollata Megathiris detruncata 13C, 4V, 14D Co tei Badenian M.62.985. Megathyris decollata Megathiris detruncata 2C Co tei Badenian M.62.986. Argyrotheca subcuneata Argyrotheca cordata

Megathiris detruncata 1C1V

Co tei Badenian

M.62.987. Argyrotheca subcordata Argyrotheca cordata 2C Co tei Badenian M.62.988. Argyrotheca subcordata Argyrotheca cordata 3C Co tei Badenian M.62.989. Crania subrostrata Megerlia truncata 1D Co tei Badenian M.62.995. Brachiopoda Megerlia truncata 1D Co tei Badenian M.62.991. Terebratula cf. hoernesi Terebratula sp. 1F Eisenstadt Badenian M.62.992. Terebratula macrescens Terebratula macrescens 1V, 1D Eisenstadt Badenian M.62.993. Megerlia margaritata Megerlia margaritata 1C Buciumi Badenian M.62.994. Terebratula sp. Terebratula sp. 2F St Margerethen Badenian M.62.996. Terebratula macrescens Terebratula macrescens 8C, 6V, 1D Eisenstadt Badenian M.62.997. Terebratula macrescens Terebratula macrescens 2C, 16V, 3D Eisenstadt Badenian M.62.998. Terebratula macrescens Terebratula macrescens 1V, 1D Eisenstadt Badenian M.62.999. Terebratula macrescens Terebratula macrescens 3C, 3V, 1D Eisenstadt Badenian M.62.1000. Terebratula macrescens Terebratula macrescens 3C, 3V, 1D Eisenstadt Badenian

BITNER & DULAI


82M.62.1001. Terebratula macrescens Terebratula macrescens 2C, 2V Eisenstadt Badenian M.62.1002. Terebratula macrescens Terebratula macrescens 1C, 3V Eisenstadt Badenian M.62.1003. Terebratula macrescens Terebratula macrescens 1F Eisenstadt Badenian M.62.1004. Terebratula macrescens Terebratula macrescens 7V Eisenstadt Badenian M.62.1005. Megathyris decollata Megathiris detruncata 1C, 1D Eisenstadt Badenian

Unpublished collection

Inventory Old name New name Material Locality Age

M.63.284. Terebratula sp. Gryphus miocaenicus 1V Salgótarján Karpatian M.63.260. Terebratula sp. Gryphus miocaenicus 2V Salgótarján Karpatian M.63.261. Terebratulina sp. ?Bivalvia - Salgótarján Karpatian M.63.264. Terebratula sp. Gryphus miocaenicus 9V Salgótarján Karpatian M.61.6167. Terebratula sp. Gryphus miocaenicus 2V Fót Karpatian M.61.6175. Terebratula sp. Gryphus miocaenicus 2C, 4V Fót Karpatian M.62.233. Terebratula sp. Terebratula sp. 2V, 1D, 6F Fót Karpatian M.61.6718. Terebratula macrescens Terebratula macrescens 1V Püspökhatvan Karpatian M.61.6717. Brachiopoda Terebratula sp. 1V, 1D Püspökhatvan Karpatian M.61.6719. Terebratula sp. Terebratula sp. 4V, 2D Püspökhatvan Karpatian M.61.6506. Terebratula sp. Terebratula sp. 1C Mogyoród Karpatian M.61.6523. Cistella sp. Gryphus miocaenicus 2C Mogyoród Karpatian M.61.6516. Cistella sp. Gryphus miocaenicus 3C, 1V, 1D Mogyoród Karpatian M.61.6522. Cistella sp. Gryphus miocaenicus 18C, 9V, 2D Mogyoród Karpatian M.61.6521. Cistella sp. Gryphus miocaenicus 22C, 7V, 4D Mogyoród Karpatian M.61.5979. Brachiopoda Terebratula sp. 1V Mátyásföld Karpatian M.62.4737. Terebratula sp. Terebratula sp. 1V Fert rákos Badenian M.60.5023. Terebratula sp. Bivalvia - Várpalota Badenian M.62.6666. Terebratula grandis Terebratula sp. 1C Sámsonháza Badenian M.62.6668. Terebratula sinuosa Terebratula sp. 1C Sámsonháza Badenian M.62.6965. Cistella sp. Platidia anomioides 1C Sámsonháza Badenian M.61.8400. Terebratula sp. Terebratula sp.

Aphelesia sp.13C, 5V 3C, 2V

Márkháza Badenian

M.61.8385. Terebratula macrescens Terebratula macrescens 6C, 3V, 1D Márkháza Badenian M.61.8411. Terebratula sinuosa Terebratula sp. 4C Márkháza Badenian M.61.8862. Miletia angulosa Terebratula sp. 1V Márkháza Badenian M.62.2721. Argyrotheca sp. Platidia anomioides 1C Márkháza Badenian M.61.8842. Argyrotheca cf. neapolitana Partly: Terebratula sp. 1C Márkháza Badenian M.61.8838. Hemithiris acuta Aphelesia sp. 1C Márkháza Badenian M.61.8844. Gryphus miocaenicus Gryphus miocaenicus 1C Márháza Badenian M.61.8452. Cistella sp. Platidia anomioides 1C Márkháza Badenian M.61.8837. Terebratula macrescens Terebratula macrescens 2C Márkháza Badenian M.61.8847. Terebratula macrescens Terebratula sp. 1F Márkháza Badenian M.62.3143. Argyrotheca neapolitana Platidia anomioides 4C, 6V, 3D Mátraverebély Badenian M.62.3144. Hemithiris acuta Aphelesia sp. 1V Mátraverebély Badenian M.62.3145. Terebratula sp. Partly: Platidia anomioides 1C Mátraverebély Badenian M.62.3051. Cistella sp. Platidia anomioides 3C, 1V Mátraverebély Badenian M.62.3142. Brachiopoda Platidia anomioides 2C, 1V Mátraverebély Badenian M.62.3879. Terebratula sp. Gryphus miocaenicus 4C Mátrasz l s Badenian M.62.3880. Terebratula sp. Gryphus miocaenicus 2V Mátrasz l s Badenian M.62.3881. Terebratula sp. Gryphus miocaenicus 1C, 1V, 1D Mátrasz l s Badenian M.62.3882. Terebratula sp. Gryphus miocaenicus 1C Mátrasz l s Badenian M.62.3883. Megathyris sp. Argyrotheca cuneata 2C Mátrasz l s Badenian M.62.3884. Argyrotheca sp. Argyrotheca cordata 2C Mátrasz l s Badenian M.62.3878. Terebratulina sp. Argyrotheca cordata 1C Mátrasz l s Badenian M.61.8343. Brachiopoda Platidia anomioides 1C Nógrádszakál Badenian M.62.4076. Terebratula macrescens Terebratula sp. 2V, 4D Eisenstadt Badenian M.59.1000. Terebratula + Serpula Terebratula sp. 1V Eisenstadt Badenian M.59.1123. Terebratula macrescens Terebratula macrescens 1C, 1D Eisenstadt Badenian M.62.4077. Terebratula macrescens Gryphus miocaenicus 1C, 1D Eisenstadt Badenian M.62.4078. Terebratula macrescens Terebratula sp. 5F Eisenstadt Badenian M.62.4079. Terebratula ampula Terebratula sp. 6F Eisenstadt Badenian M.62.4653. Terebratula sp. Terebratula sp. 1V Leitha Mts Badenian M.65.393. Terebratula sp. Terebratula sp. 1C Horodenka Badenian M.65.392. Terebratula grandis Terebratula sp. 1V Horodenka Badenian M.60.4570. Terebratula sp. Terebratula sp. 1C, 3V Horodenka Badenian M.60.4581. Terebratula sp. Terebratula sp. 1V Horodenka Badenian M.60.4558. Terebratula grandis Terebratula sp. 1C Horodenka Badenian M.60.9216. Terebratula grandis Terebratula sp. 5C, 2V Lopadea Vache Badenian M.60.9046. Terebratula sp. Terebratula sp. 1C, 1D Lopadea Vache Badenian M.60.10002. Crania abnormis Not brachiopod - Co tei Badenian

BITNER, M. A. & DULAI, A. (2004): Revision of Miocene brachiopods of the Hungarian Natural History...

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Transcript of BITNER, M. A. & DULAI, A. (2004): Revision of Miocene brachiopods of the Hungarian Natural History...