2016-2017 ANNUAL REPORT - Marine Biological Laboratory

2016 -2017 MBL ANNUAL REPORT i1

2016-2017 ANNUAL REPORT

i2 MBL ANNUAL REPORT 2016 -2017

Contents

1 Message from Interim Co-Directors

2 Highlights

6 Financials

9 Gifts

22 Leadership

23 Advisory Bodies

24 Scientists at MBL

The MBL Annual Report is published by the Marine Biological Laboratory. Although the greatest possible care has been taken in the preparation of this record, the MBL recognizes the possibility of inaccuracies. If any are noted, please accept our apology and advise us of any corrections to be made.

Office of CommunicationsMarine Biological Laboratory7 MBL StreetWoods Hole, MA [email protected]

2016 -2017 MBL ANNUAL REPORT 1

In May 2017 we were privileged to add the role of Interim Co-Directors to our existing long-time connections to the Marine Biological Laboratory.

As we started our work, a primary goal was to listen to the diverse perspectives of the MBL community while a search was conducted for long-term leadership. Through the interim year we worked to implement strategic initiatives for the MBL that had been developed over the past four years. The research and education highlights shared in this report reflect some of the progress made in these initiatives over the 2016-2017 time period.

The MBL’s transformative impact on science is a result of the ways in which it has fostered collaboration and innovation for well over a century. Throughout this time, the critical asset of the MBL has been, and continues to be, its people—the scientists, staff and diverse supporters that make its distinctive research and learning possible. In that regard, we thank you for your input, energy, and engagement as we support and champion the MBL in our new roles as Interim Co-Directors.

Sincerely,

From the

President and

Director

Melina Hale, Vice Provost for Academic Initiatives, and William Rainey Harper Professor in the College and the Department of Organismal Biology and Anatomy, University of Chicago

Neil Shubin, Robert R. Bensley Professor in the Department of Organismal Biology and Anatomy, and Associate Dean for Academic Strategy in the Biological Sciences Division

2 MBL ANNUAL REPORT 2016 -2017

“Smart” Cephalopods Trade Off Genome Evolution for Prolific RNA Editing

A study led by Senior Scientist Joshua Rosenthal suggested that the cephalopod’s evolutionary path to neural sophistication includes a novel mechanism: prolific RNA editing at the expense of evolution in their genomic DNA. The story was widely covered in the popular media, including the New York Times and The Atlantic. Rosenthal is a neurobiologist whose research focuses on developing new, genetically tractable marine model organisms and understanding how they are affected by a changing environment. He is leading the MBL’s Marine Organisms strategic theme, bringing cutting edge imaging, genomic, and computational tools to bear on the use of marine organisms to better understand basic biology. Specifically, Rosenthal and colleagues have embarked on a groundbreaking effort to culture cephalopods in the laboratory with the goal of creating a new genetic model system.

Four Scientists are Named Distinguished Scientists

Four scientists were named Distinguished Scientists of the Marine Biological Laboratory. The title is conferred to highly accomplished scientists whose exceptional work at the Marine Biological Laboratory is of recognized national and international significance. The honors were conferred at a meeting of the MBL Society. Following the announcement, a permanent exhibit honoring the Laboratory’s Distinguished Scientists was unveiled in the lobby of Lillie Building. Named Distinguished Scientists of the Marine Biological Laboratory were John E. Dowling, Avram Hershko, Joan V. Ruderman, and Ronald D. Vale. Previously named MBL Distinguished Scientists are: Shinya Inoué (1986); John E. Hobbie (2005); Osamu Shimomura (2008); Jerry M. Melillo (2010); Mitchell L. Sogin (2014); and Ivan Valiela (2014).

David Mark Welch, Jennifer Morgan Receive Appointments

Evolutionary biologist David Mark Welch, Director of the Josephine Bay Paul Center and MBL Senior Scientist was named Interim Director, Division of Research. Mark Welch has been associated with the MBL for more than 15 years. He has been Director of the Bay Paul Center since 2013. MBL Associate Scientist Jennifer Morgan was named Director of the Eugene Bell Center for Regenerative Biology and Tissue Engineering. Morgan was recruited to the MBL in 2012 and has served as the Associate Director of the Bell Center since that time. Morgan’s research program focuses on understanding how synapses normally function and how synaptic function is affected by neurological diseases. They succeed Jonathan Gitlin, who retired in June 2017, and previously held both positions.

Highlights


Long-term Ecological Research Sites Receive Grant Renewals

Anne Giblin, Ecosystems Center Interim Director was awarded a $6.8 million National Science Foundation grant to study how sea-level rise, climate change, and human impacts are reshaping the marshes and estuaries behind Plum Island, Mass., the largest remaining marsh area in the northeastern United States. Giblin received the grant as lead investigator of NSF’s Long-Term Ecological Research (LTER) program at Plum Island Sound. MBL ecologists have been conducting research at the site since the late 1980s and have directed the Plum Island LTER since its creation in 1998. For the fourth consecutive decade, MBL scientists will lead the nation’s primary research project on Arctic ecology and its responses to climate warming, with $6.8 million in NSF funding awarded to Senior Scientist Edward B. Rastetter, lead investigator of the LTER site at Toolik Lake, Alaska.

Hibbitt Early Career Fellowship Program Established

Three talented postdoctoral scientists were named the inaugural Hibbitt Early Career Fellows in Spring 2017. The fellowships were awarded to Caroline Albertin, Kristen Hunter-Cevera, and Duygu Özpolat. The opportunity attracted a strong applicant pool, and 100% of the offers made were accepted. Albertin is interested in the unusual body plans of the cephalopods and was part of a team that reported the first genome sequence from a cephalopod. Hunter-Cevera studies the population dynamics of marine cyanobacterium and how changes in ocean temperature affect key species of phytoplankton. Özpolat is a developmental biologist who studies the mechanisms of reproductive cell regeneration. The program, funded by a generous gift from Susan and David Hibbitt, MBL Trustee, provides talented scientists the chance to set up research programs of their own as an alternative to traditional postdoctoral positions.

Education Program Score High Marks from Alumni; Launches New Course

The MBL was exceedingly highly rated in an independent survey of MBL Discovery Course alumni from 2005-2016. The overwhelming majority of alumni respondents felt their time at MBL was well spent—even extraordinary in many cases. 99% of the more than 600 respondents rated it as either very worthwhile or transformative. In 2016 a new course titled Physical Biology of the Cell, directed by Rob Phillips (Caltech) and Hernan Garcia (UCSF), explored a broad array of topics from modern biology using the language of physics and mathematics. The course was aimed at those interested in learning more about how to construct theoretical models of biological systems as well as the use of computational tools to simulate and test the predictions of those models.


Ron Vale Receives Shaw Prize for Discovery of Motor Protein Family, Kinesin, at MBL

Ronald Vale, who led a group of scientists at the MBL to the seminal discovery of kinesin, one of the major families of motor proteins, was awarded the prestigious 2017 Shaw Prize in Life Science and Medicine, often called the “Nobel of the East.” Vale was a student visiting the MBL in the mid-1980s when he and collaborators purified a mystery motor protein from the axoplasm of squid, which they later called kinesin. The discovery opened up a still-flourishing research field exploring kinesin’s critical biological functions including moving molecular cargo in cells and separating chromosomes during cell division. Vale is a Professor at the University of California-San Francisco, a Howard Hughes Medical Institute Investigator, and MBL Distinguished Scientist. He received the Shaw Prize along with Ian Gibbons of University of California-Berkeley, who discovered the other major family of motor proteins, dynein, in the 1960s.

National Xenopus Resource Expands; Provides Critical Resources to Scientists

The National Xenopus Resource (NXR) provides customized transgenic and mutant frogs to the Xenopus research community, as well as advanced training workshops and research facilities for visiting scientists. With growth over the last six years and advances in genome technology leading to new mutant lines of animals, the need for expansion was identified. Marko Horb, NXR Director and Principal Investigator received an NIH expansion grant that funded an expansion that will allow the facility to hold 100-200 additional lines and keep pace with the changing technologies. The new space will also help the NXR to continue to attract visiting scientists.

Two New Microscopes Developed at MBL

Taking a cue from medical imaging, scientists working in the Whitman Center invented a multi-view microscope that captures higher-resolution, 3D images of live cells and tissues without upping the dose of potentially harmful radiation the specimens receive. MBL Fellows Hari Shroff (National Institute of Biomedical Imaging and Bioengineering) and Patrick La Rivière (University of Chicago) reported that the new system achieved resolution which is double the volumetric resolution of traditional fluorescence microscopy methods. Another new microscope, developed by MBL scientists Tomomi Tani and Shalin Mehta (now at Chan Zuckerberg Biohub) can track the position and orientation of individual molecules in living cells—nanoscale measurements that until now have posed a significant challenge. Tani and Mehta used the microscope to address various biological questions in collaboration with a team of MBL Resident and Whitman Scientists.


Financials

Fiscal Year

June 30, 2017

ENDOWMENT VALUE

68.7

79.0

82.1

76.3

80.2

60

65

70

75

80

85

June 30, 2013

June 30, 2014

June 30, 2015

June 30, 2016

June 30, 2017

$

$

$

$

$

$

OPERATING REVENUE OF $39.6 MILLION

OPERATING EXPENSES OF $42.8 MILLION

Federal Grants 32%

Resident Research 56%

Education 24%

Visiting Research

9%

Non-federal Grants 12%

Gifts 9%

Fees for Service

11%

Endowment Income

11%

Income from Affiliates

21%

Other Revenue 4% Other Programs 5%

Conferences, Housing and Dining 6%

The financial statements of the Marine Biological Laboratory for the fiscal year ending June 30, 2017 were audited by KPMG. Complete financial statements are available on request from: Financial ServicesMarine Biological Laboratory7 MBL StreetWoods Hole, MA 02543

$

$

$

$

$

in m

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I am pleased to report on the Marine Biological Laboratory’s development progress for the 2017 Fiscal Year: July 1, 2016 through June 30, 2017.

In FY17 we raised $7,076,738 in new private support for MBL’s research and educational programs from over 900 individuals, foundations, and companies. This impressive figure is a testament to the MBL’s history of scientific excellence, our powerful impact on biological research, and the vision of an equally impactful and sustainable future.

Funds raised during this time include several major gifts in support of MBL programs and initiatives, including: David and Susan Hibbitt, who provided $2 million to establish the Hibbitt Early Career Fellows Program, giving talented scientists the chance to establish their own research programs; The Burroughs Wellcome Fund, who pledged $660,000 in support of the MBL Physiology and Embryology courses; and The Grass Foundation, who pledged $486,000 in support of the Neurobiology and Neural Systems and Behavior courses. Please see a full list of our supporters in the pages that follow.

We also raised $1,008,966 in Annual Giving in FY17 — unrestricted funds that are immediately available to meet operational needs. Much of this impressive figure is made up of gifts of $50, $20, and sometimes even $10, but they can add up to a large sum. We continued to expand annual giving, participating in our second annual ‘Giving Day,’ a joint fundraising effort with the University of Chicago. This one-day event added $46,000 to our annual fund, a wonderful group effort by the development teams at MBL and the University.

The MBL community is wide-reaching, and there is strength in numbers. Every gift, whether it’s $20 or $20,000, makes an impact. I’m grateful to our Trustees, MBL Council and Society members, scientists, alumni, faculty, friends, and to every one of our extraordinary donors.

On behalf of the Development Committee, the Board of Trustees, and the staff of the MBL – thank you all for your generosity!

Gratefully, Jeffery Pierce, Chair

Report of the Development Committee


Gifts The Marine Biological Laboratory gratefully acknowledges the generous contributions and pledges of the individuals, foundations, and companies who supported our programs and operations.

MAJOR GIFTS

$1,000,000 + David and Susan Hibbitt

$100,000 - $999,999Anonymous (3)Burroughs Wellcome FundThe Grass FoundationGeorge Frederick Jewett Foundation

EastJames S. McDonnell FoundationOwens Family FoundationThe E. Joel Safran TrustSimons FoundationThe G. Unger Vetlesen Foundation

RESTRICTED GIFTS

$50,000 - $99,999Anonymous (4)International Brain Research

Organization (IBRO)The Estate of Louise M. LuckenbillMerle Mizell

$10,000 - $49,999Anonymous (2)The Agouron InstituteThe Foundation for Agricultural

IntegrityAmerican Chemistry CouncilMargaret C.** and Francis P. BowlesThe Goldschmidt FoundationNikon Instruments, Inc.Ann E.** Stuart and John W.** MooreWilliam Townsend Porter FoundationEdwin S. Webster Foundation

$5,000 - $9,999Animal Welfare InstituteThe Company of BiologistsThe Kavli FoundationJack** and Francine LevinMarch of Dimes FoundationSociety for the Study of ReproductionJohn N.* and Karen Vournakis In memory of David T. Sullivan

$1,000 - $4,999Anonymous (1)Constance M. and Dean C. AllardPorter W. Anderson** and Hortense GerardoAquaneering, Inc.Arivis AGPeter B.** and Margaret T. ArmstrongDon and Dee AukampThe Baila FoundationMolly Bang and Jim GreenPhilip Batterham In memory of David T. SullivanNew England BioLabs, Inc.Gloria S. Borgese In memory of Thomas A. BorgeseM. Kathryn S. and Thomas A. Brown In memory of John W. Saunders, Jr.Captain Kidd Restaurant In memory of William CrowleyPaul N. ChervinFrank M.** and Julia S. ChildJohn D. Clayton and Will Jolley In memory of John W. Saunders, Jr.David and Patricia Cohen In memory of William D. CohenMarion Cohen In memory of William D. CohenDevelopmental DynamicsEuFishBioMedPamela and David FollettRichard Harland In memory of John W. Saunders, Jr.William Allen Held In memory of John W. Saunders, Jr.Iwaki America

William Jeffery and Meredith Yeager-Jeffery

In memory of John W. Saunders, Jr.Jack and Pam Jolley In memory of John W. Saunders, Jr.Paul B. LazarowJason and Jennifer Meyers In memory of John W. Saunders, Jr.Nature MethodsOrganization for Computational

NeurosciencesPentair Aquatic HabitatsSarah Elizabeth S. Reeder In memory of John W. Saunders, Jr.Patricia** and Charles RobertsonJohn M. and Donna K. RussellWendy C. SalmonJack W. and Lorie Saunders In memory of John W. Saunders, Jr.Robert D. Schatz In honor of Annie SchatzSociety of General PhysiologistsMarie and David Sullivan In memory of David T. SullivanTechnic, Inc.Dyann** and Peter K. WirthYingzi Yang In memory of John W. Saunders, Jr.

$1 - $999Anonymous (6)Sharon L. Amacher In memory of John W. Saunders, Jr.American Society for MicrobiologyRobert and Elizabeth Angelici In memory of Gwen P. Szent-GyörgyiApplied Scientific Instrumentation, Inc.Ricardo C. AranedaBobbi Bailin In memory of Gwen P. Szent-GyörgyiSpencer L. Baird and Judy P. Spencer In memory of Gwen P. Szent-GyörgyiNancy Barnett In honor of Douglas BurrillBlueline NDT-LLCGiovanna Breu


Melinda Browne In memory of John J. ValoisRobert and Linda Campbell In memory of Gwen P. Szent-GyörgyiChenbei Chang In memory of John W. Saunders, Jr.Jian-Fu Chen In memory of John W. Saunders, Jr.Ida Chow In memory of John W. Saunders, Jr.David ClarkeJohn R. and Annette W. Coleman In memory of John W. Saunders, Jr.Mary E. Dickinson In memory of John W. Saunders, Jr.Colleen DicksonAlberto Rosello Diez In memory of John W. Saunders, Jr.Gregg Duester In memory of John W. Saunders, Jr.Walid Fakhouri In memory of John W. Saunders, Jr.Kathryn Fowler In memory of Andrew Szent-Györgyi

and Gwen P. Szent-GyörgyiScott E. Fraser In memory of John W. Saunders, Jr.Anne FreemanJeffrey GardnerMark Gasarch In honor of Eve GasarchMargaret A. Geist In memory of John W. Saunders, Jr.Gene Tools, LLCHarmony Glazebrook-PittetGustav Grosch and Jane KuleszaJeffrey M. Gross In memory of John W. Saunders, Jr.Donald and Donna Gruol In memory of Gwen P. Szent-GyörgyiDiane B. Gumz In memory of Gwen P. Szent-GyörgyiClaudia Y. Hansen In memory of Gwen P. Szent-GyörgyiBrian D. Harfe In memory of John W. Saunders, Jr.

Joseph HeitmanEllen HertzmarkRichard HoltIntavis Inc.International Zebrafish SocietyDavid Isenberg and Paula BlumenthalLorette Javois In memory of John W. Saunders, Jr.Jane Johnson In memory of John W. Saunders, Jr.Barbara Woll Jones In memory of Robert C. KingFreda Kaminer In memory of Gwen P. Szent-GyörgyiLouie M. and Brenda Kerr In memory of John W. Saunders, Jr.Robert Kosher and Caroline Dealy In memory of John W. Saunders, Jr.Robert E. Krumlauf In memory of John W. Saunders, Jr.Mark Lewandoski In memory of John W. Saunders, Jr.Huili Li In memory of John W. Saunders, Jr.Aimin Liu In memory of John W. Saunders, Jr.Thomas E. LovejoyBob Maccaferri In memory of Gwen P. Szent-GyörgyiSusan Mackem In memory of John W. Saunders, Jr.Anthony P. Mahowald In memory of John W. Saunders, Jr.Anne F. Martin In memory of Gwen P. Szent-GyörgyiDavid R. McClay and N. Allison Haltom In memory of John W. Saunders, Jr.Andrew McMahon In memory of John W. Saunders, Jr.David A. and Virginia I. MillerSusan MorseMary C. Mullins In memory of John W. Saunders, Jr.Elizabeth Myer In memory of Gwen P. Szent-Györgyi

Lee A. Niswander In memory of John W. Saunders, Jr.Michael Norton In memory of Catherine N. NortonElizabeth PennisiHoward Prior and Molly Aston In memory of Gwen P. Szent-GyörgyiFred and Gloria RhoadesTania J. Rozario In memory of John W. Saunders, Jr.Martha Jane Salyers In memory of Abigail SalyersJoseph W.** and Jean M. Sanger In memory of John W. Saunders, Jr.Richard A. Schneider In memory of John W. Saunders, Jr.Michael D. Shapiro In memory of John W. Saunders, Jr.Noriko Shimizu and Courtland Yockey In memory of Gwen P. Szent-GyörgyiStanley Shostak In memory of John W. Saunders, Jr.Ralph Slepecky In memory of David T. SullivanBradley and Kirsten Smith In memory of Gwen P. Szent-GyörgyiPaul F. Smith, Jr. In memory of Gwen P. Szent-GyörgyiMitchell L. Sogin and Laurel MillerHeather Southwick In memory of Gwen P. Szent-GyörgyiKathy A. Suprenant and William L.

DentlerSutter InstrumentsGerard L. and Mary SwopeBen G. Szaro In memory of John W. Saunders, Jr.Alexis Tapanes-Castillo In memory of John W. Saunders, Jr.Tecniplast USA, Inc.Jane K. Thompson In memory of Gwen P. Szent-GyörgyiPaul and Clair Tracy In memory of Gwen P. Szent-GyörgyiDavid C. Turner In memory of David T. Sullivan

United States Biological CorporationFrederica W. Valois In memory of Gwen P. Szent-GyörgyiSteven Vokes In memory of John W. Saunders, Jr.William W. WalkerMs. John B. Wallingford In memory of John W. Saunders, Jr.Leonard and Eve WarrenScott Weatherbe In memory of John W. Saunders, Jr.Janis C. Weeks and William M. RobertsRonald Zweig and Christina Rawley

GIFTS IN KINDFine Science ToolsHuntington** and Vicki Willard


GIFTS AND GRANTS TO FELLOWSHIPS, SCHOLARSHIPS, AND OTHER FUNDSWe gratefully acknowledge donors to the following funds that enable the Marine Biological Laboratory to stay competitive by attracting and supporting top researchers and students.

Frederik B. and Betsy G. Bang Research AwardsMolly Bang and Jim GreenJack** and Francine Levin

The Josephine Bay Paul Center for Comparative Molecular Biology and EvolutionThe G. Unger Vetlesen Foundation

Biology of Parasitism CourseAnonymous (1)Dyann** and Peter K. Wirth

Bernard Davis Scholarship FundPorter W. Anderson** and Hortense

Gerardo

Thomas A. and Gloria S. Borgese Rare Book and Journal FundAnonymous (1)Gloria S. Borgese In memory of Thomas A. Borgese

Brown University and MBL Partnership ProgramRichard Holt

Central Microscopy Facility SupportAnonymous (1)

Chief Operating Officer Discretionary FundGeorge Frederick Jewett Foundation East

William D. Cohen Endowed Fund for Undergraduate EducationDavid and Patricia Cohen In memory of William D. CohenMarion Cohen In memory of William D. Cohen

Computational Image Analysis CourseArivis AGNature Methods

Philip Dunham Memorial Fund In memory of Philip DunhamFrank M.** and Julia S. ChildJohn M. and Donna K. RussellRalph Slepecky

Eugene Floyd DuBois Memorial FundHarmony Glazebrook-Pittet

Ecosystems Long-Term Ecological Research Projects FundAnonymous

The Mac V. Edds, Jr. Endowed Scholarship FundLouise M. Luckenbill and Harvey Culbert

Embryology Course Educational GrantBurroughs Wellcome Fund

Embryology Course ScholarshipThe Company of Biologists LimitedMarch of Dimes Foundation

Falmouth Forum EndowmentAnonymous (2)Nancy Barnett In honor of Douglas BurrillBarbara Woll Jones In memory of Robert C. KingRonald Zweig and Christina Rawley

Send a Falmouth Teacher to Toolik, AlaskaAnonymous (1)Don and Dee Aukamp

Margaret C.** and Francis P. BowlesPamela and David FollettDavid and Susan HibbittSusan MorsePatricia** and Charles RobertsonGerard L. and Mary Swope

Frontiers in Reproduction Course Education GrantBurroughs Wellcome Fund

Frontiers in Reproduction Course ScholarshipAnonymous (1)Society for the Study of Reproduction

Daniel S. and Edith T. Grosch Scholarship FundGustav Grosch and Jane Kulesza

Aline D. Gross ScholarshipFreda KaminerTechnic, Inc.

Roger Hanlon Discretionary FundGeorge Frederick Jewett Foundation East

Tay Hayashi Lectureship in Cell PhysiologyFreda Kaminer

Pamela Clapp Hinkle Community FundPorter W. Anderson** and

Hortense Gerardo

Pamela Hinkle Communications Internship in Digital EngagementMargaret C.** and Francis P. Bowles

History of Biology ProgramEdwin S. Webster Foundation

The John E. Hobbie Fund for ScienceDavid ClarkeThomas E. Lovejoy

IBRO Neuroscience ScholarshipsInternational Brain Research

Organization (IBRO)

Shinya Inoue Endowed Lecture in AQLMWendy C. SalmonJoseph W.** and Jean M. Sanger

Irvin Isenberg LectureDavid Isenberg and Paula BlumenthalFreda Kaminer

Benjamin Kaminer ScholarshipFreda Kaminer

Jane Lazarow Stetten Scholarship FundPaul B. Lazarow

Logan Science Journalism ProgramGiovanna BreuElizabeth Pennisi

Marine Resources Center Discovery in Marine OrganismsAnonymous (1)Animal Welfare Institute

Marine Resources Center Infrastructure EnhancementsAnonymous (1)

The MBL Hibbitt Early Career FellowsAnonymous (1)David and Susan Hibbitt

MBLWHOI Library FundLeonard and Eve Warren

MBL Library Planning and Renovation FundMerle Mizell

Methods in Computational Neuroscience Course ScholarshipOrganization for Computational

Neurosciences


Microbial Diversity CourseSimons Foundation

Microbial Diversity Course Geomicrobiology SymposiumThe Agouron Institute

Microbial Diversity Course ScholarshipThe Goldschmidt Foundation

James A. and Faith Miller Research AwardsDavid A. and Virginia I. Miller

Mobile Genetic Elements WorkshopNew England BioLabs, Inc.

Molecular Mycology CourseJoseph Heitman

Monitoring Greenhouse Gas Fluxes and Crop Photosynthesis ProjectThe Foundation for Agricultural Integrity

John W. Moore and Ann E. Stuart Endowed FundAnn E.** and John W.** Moore

Mountain Memorial FundConstance M. and Dean C. Allard

Neural Systems and Behavior and Neurobiology Courses Education GrantThe Grass Foundation

Neural Systems and Behavior Course Scholarship Endowed FundJanis C. Weeks and William M. Roberts

Neurobiology CourseRicardo C. Araneda

Nikon Research AwardsNikon Instruments, Inc.

Catherine N. Norton FellowshipCaptain Kidd Restaurant In memory of William CowleyColleen DicksonMichael Norton In memory of Catherine N. Norton

Physiology Course Education GrantBurroughs Wellcome Fund

Physiology CourseKathy A. Suprenant and William L. Dentler

Physiology Course EndowmentJoseph W.** and Jean M. Sanger

Plastic Marine Debris Fragmentation, Density, and Deposition ProjectAmerican Chemistry Council

Putting History and Philosophy of Life Sciences to Work Research ProjectJames S. McDonnell Foundation

William Ratzan Research SupportAnonymous (1)

Joshua Rosenthal Laboratory SupportOwens Family Foundation

S. O. Mast Endowed ScholarshipFred and Gloria Rhoades

Abigail Salyers Endowed Scholarship in Microbial DiversityAmerican Society for MicrobiologyJeffrey GardnerMartha Jane Salyers In memory of Abigail SalyersMitchell L. Sogin and Laurel Miller

John Saunders Memorial Fund In memory of John W. Saunders, Jr.Anonymous (4)Sharon L. AmacherM. Kathryn S. and Thomas A. Brown

Chenbei ChangJian-Fu ChenIda ChowJohn D. Clayton and Will JolleyJohn R. and Annette W. ColemanMary E. DickinsonGregg DuesterWalid FakhouriScott E. FraserMargaret A. GeistJeffrey M. GrossBrian D. HarfeRichard HarlandWilliam Allen HeldLorette JavoisWilliam Jeffery and Meredith

Yeager-JefferyJane JohnsonJack and Pam JolleyLouie M. and Brenda KerrRobert Kosher and Caroline DealyRobert E. KrumlaufMark LewandoskiHuili LiAimin LiuSusan MackemAnthony P. MahowaldDavid R. McClay and N. Allison HaltomAndrew McMahonJason and Jennifer MeyersMary C. MullinsLee A. NiswanderSarah Elizabeth S. ReederAlberto Rosellό-DiezTania J. RozarioJoseph W.** and Jean M. SangerJack W. and Lorie SaundersRichard A. SchneiderMichael D. ShapiroStanley ShostakBen G. SzaroAlexis Tapanes-CastilloSteven VokesMs. John B. WallingfordScott WeatherbeYingzi Yang

Semester Environmental Science Program SupportAnonymous (1)The Baila FoundationAnne FreemanMark Gasarch In honor of Eve GasarchEllen HertzmarkRobert D. Schatz In honor of Annie SchatzWilliam W. Walker

Society of General Physiologists ScholarshipSociety of General Physiologists

SPINES ScholarshipWilliam Townsend Porter Foundation

David Sullivan Memorial Fund In memory of David T. SullivanPhilip BatterhamRalph SlepeckyMarie and David SullivanDavid C. TurnerJohn N.* and Karen Vournakis

Andrew Szent-Györgyi Endowed Lecture in PhysiologyKathryn Fowler In memory of Andrew Szent-Györgyi

and Gwen P. Szent-GyörgyiFreda Kaminer

Eva Szent-Györgyi Scholarship Fund In memory of Gwen P. Szent-GyörgyiRobert and Elizabeth AngeliciBobbi BailinSpencer L. Baird and Judy P. SpencerRobert and Linda CampbellMrs. Kathryn FowlerDonald and Donna GruolDiane B. GumzClaudia Y. HansenFreda KaminerBob Maccaferri


Anne F. MartinElizabeth MyerHoward Prior and Molly AstonNoriko Shimizu and Courtland YockeyBradley and Kirsten SmithPaul F. Smith, Jr.Heather SouthwickJane K. ThompsonPaul and Clair TracyFrederica W. Valois

Undergraduate Research Programs for Underrepresented MinoritiesBurroughs Wellcome Fund

John J. Valois Marine Resources Center EndowmentAnonymous (1)Melinda Browne In memory of John J. ValoisFreda Kaminer

Whitman Center General EndowmentPeter B.** and Margaret T. Armstrong

Walter L. Wilson Endowed Scholarship FundPaul N. Chervin

Women in Ecological Science FundAnonymous (1)

Zebrafish Development and Genetics CoursePentair Aquatic Habitats

Zebrafish Development and Genetics Course 20th Anniversary SymposiumApplied Scientific Instrumentation, Inc.Aquaneering, Inc.Blueline NDT-LLCThe Company of Biologists LimitedDevelopmental DynamicsEuFishBioMedGene Tools, LLCIntavis Inc.

International Zebrafish SocietyIwaki AmericaThe Kavli FoundationSutter InstrumentsTecniplast USA, Inc.United States Biological Corporation

GIFTS TO THE MARINE BIOLOGICAL LABORATORY ANNUAL FUNDThrough their annual unrestricted gifts, annual fund donors enable the Marine Biological Laboratory to cover costs not met by grants or fees, seed key strategic initiatives, and provide important support to its scientists and programs.

The Chairman’s Circle $25,000 or moreDavid and Susan HibbittOwens Family FoundationThe Pannell Family Charitable FundJeffrey and Margaret PierceKathy and George PutnamLeslie J.* and Elizabeth M.* WilsonRobert J. Zimmer

The Director’s Circle $10,000 - $24,999Anonymous (4) In memory of Thomas Clowes (1)Alison and Robert AmentPorter W. Anderson** and Hortense GerardoMartha W.** and William C. CoxDavid FithianMargaret and Timothy FofonoffPatrick Gage and Irina Wilson GageThomas F. and Virginia C. GreggWilliam and Lauren HuyettEric D. IsaacsChristopher G. KennedyEthan A. and Lisa LernerAnna LysakowskiMarianne F. MilkmanRobert W. and Carroll C. D. PierceTimothy Springer and Chafen LuCarolyn and John StimpsonHuntington and Vicki Willard

The President’s Club $5,000 - $9,999Anonymous (1)Don and Dee AukampDavid and Sandra Bakalar

John and Lisa BatterJoshua and Lisa BernsteinNorman** and Diane BernsteinMarianne and Eric BillingsPhyllis S. BockEd and Amy BrakemanAdam de Sola Pool, Kristina M. Gjerde,

and Darius PoolChristopher D. EarlMichael N. Fenlon and Linda SallopJean-François Formela and Rachel

Somer FormelaCraig and Nancy GibsonAlfred L. and Joan H. GoldbergWalter E. and Shirley A. MasseyJohn and Judy McCarterJeanne Guillemin-Meselson and

Matthew MeselsonBarbara Woll Jones In memory of Arthur M. SussmanBrian and Patricia KeckSaul KurlatJennifer LetitiaJoseph S. Levine and Steven A. CadwellMonique Liuzzi In memory of Anthony LiuzziGeorge** and Harmon Logan In honor of Gary G. BorisyDale and Edward J. MathiasJohn W. and Judy McCarterNikon Instruments, Inc.In memory of Michael W. DavidsonDavid and Sara E. SchnellJohn D. Sigel and Sally C. ReidRichard and Joanne SpillaneThe Seth Sprague Education &

Charitable FoundationMichael and Jill StanskyDusan Stefoski and Craig SavageGerard and Mary SwopeJohn F. SwopeFrederica W. ValoisJohn N.* and Karen VournakisRusty and Susan WarrenEarl H. WeidnerBarbara Wu and Eric Larson


Carl Zeiss Microscopy, LLC In memory of Michael W. Davidson

The Lillie Society $2,500 - $4,999Robert W.** and Jean Ashton In memory of Peter N. KivyColleen M. Cavanaugh** and Philip M.

GschwendJames N. and Nancy GallowayCharles M. GansonDavid A. and Carolyn GreeneMary Greer**Gary and Susan JacobsonPeter N. Kivy* and Joan PearlmanLaurie J. Landeau** and Robert J. MazeAnna L. and Tom LawsonMichele A. Lorand and Paul Ringel In memory of Joyce LorandThomas D.** and Patricia S. PollardPatricia** and Charles RobertsonLeonard and Eve WarrenGerald** and Ann WeissmannGary M. WesselNancy Wolf and Aric Greenfield

The Whitman Society Life MembersFrank M.** and Julie S. ChildMargaret Clowes*William Jeffrey and Meredith

Yeager-JeffreyHans Laufer

The Whitman Society $1,200 - $2,499Louise AdlerGarland E. Allen**Samuel C. and Elizabeth F. Armstrong In memory of Philip B. and Louise S.

ArmstrongDavid Baltimore and Alice S. HuangPatricia Ann BarlowGeorge and Harriet BerkowitzJoAnn Buchanan and Stephen J. Smith In memory of Christopher Vale

R. Andrew and Barbara CameronMary ClutterAdam E. CohenRita R.** and Jack H. ColwellMolly N. CornellThomas S.** and Geraldine CraneStephen and Beth CrockerPaul J.** and Maria S. De WeerJohn E.** and Judith F. DowlingJoseph and Sarah DowlingMartha S. FergusonDavid C. and Patricia S. GadsbySallie A. GiffenRobert D.** and Anne E. GoldmanTimothy** H. and Mary Helen M.

GoldsmithPhilip Grant**William H. and Margaret J. GreerRobert** and Margot HaselkornElizabeth HealdDiane E. HeckJohn G. Hildebrand** and Gail D. BurdTimothy T. HiltonPamela C. and Gregory J. HinkleJohn P. Kendall**A. Sidney and Alice KnowlesHans** and Donna Kornberg In memory of J. Woodland HastingsGeorge M.** and Sylvia T. LangfordLeonard and Ruth Ann LasterWilliam W. and Noelle LockeIrving M. and Huguette LondonLaszlo** Lorand*William K. and Winnie C. MackeyCraig C. MalbonTimothy Mitchison and Christine Marie

FieldMerle MizellJames F. and Lisa M. MooneyJudith and Thoru PedersonBertha PersonJack and Rosalyn RosenbluthEdward D. and Nancy M. SalmonOsamu and Akemi ShimomuraL. David SibleyCorinne Steel

Marjorie E. Steinberg In memory of Malcolm S. SteinbergRaymond E. StephensDorothy A. Stracher In memory of Alfred StracherDonata O. and William M. SugdenEdwin and Heather TaylorLeslie B. Vosshall and Kevin J. Lee In memory of Philip B. DunhamScott WayneAlfred M. Zeien **

Annual Fund $500 - $1,199Anonymous (1)Clay** M. Armstrong and Clara Franzini-

ArmstrongGloria S. BorgeseMargaret C.** and Francis P. BowlesAndrew and Ellen ChienMary and Jonathan ConradErnest F. and Keiko CouchCynthia and John CsernanskyGiuseppe and Mirella V. D’AlessioRobert E. and Joanne FallonProsser Gifford**Charlotte O. GoodwinKatherine Graubard and William H. CalvinPaul R. Gross**Roger T. HanlonWilliam A. HaskinsHoward H. HiattKenneth Holden and Frances C. McGuireSybil Holtzer*Shinya** and Sylvia InoueDarcy B. Kelley**Alexey L. KhodjakovEdward A.** and Kathryn F. KravitzMichael KufflerKiyoshi KusanoBirgit Rose Loewenstein** In memory of Werner R. LoewensteinDavid and Jessica Mark WelchEd and Amelia McCleskeyRichard B. McElveinMichael and Agnese Meehan

Dennis E. MeissJeanne M. NerbonneRae NishiDana T. Nojima In honor of Richard W. LinckWilliam and Sarah Odenkirk In memory of William D. CohenThe O’Herron Family FoundationRudolf and Nannette D. OldenbourgRobert H. and Pamela D. PelletreauHelen and David Piwnica-WormsKeith and Chiye PurpuraMichael and Diane RabinowitzThomas S. Reese** In memory of Christopher ValePaul B. RizzoliWilliam N. Ross and Nechama Lasser-RossJoan V.** and Gerald S. RudermanMichael G. Ryan and Linda A. JoyceRobert D. SchatzRyan Schenk In memory of Catherine N. NortonAmy R. Segal and Andrew BlumRosalind A. SegalCecily Cannan SelbyMark D. Severs In memory of Peter N. Kivy In honor of Joan PearlmanHarriet S. and Howard E. Shapiro In memory of The Sturtevants and

The LancefieldsAlexandra Shea ColinBonnie W. SimonPaul and Susan SpeerJohn H. TietjenJohn Tochko and Christina Myles-TochkoRonald D. Vale and Karen DellRichard and Dorothy VerneySally and Scott WoodsJeffrey A. and Johanna A. Zinn In memory of Donald and Margery Zinn

$100 - $499Anonymous (2)Laurence and Catherine AbbottDonald A. and Barbara R. Abt


Robin W. AckroydMelina A. AgostoJudy A. Akers In memory of Ann E. KammerDouglas G. and Kathleen Alexander In memory of Eugene O. FamilyNina Stromgren Allen**Linda Amaral and Erik R. ZettlerMichael and Marie AngeliniIrina ArkhipovaDavid S. and Nancy C. BabinHope Lincolin BakerNelson R. BartonAndre Pierre BedardDeborah Y. Beers-JonesTed Begenisich and Sherrill SpiresMarlene and George BelfortThomas L. and Mary Jo BenjaminJoan M. BernhardStephen H. and C. Lynn BishopRonald H. BlackmonThomas and Laura Bleck In memory of Frank MorrellEdward M. BlumenthalThomas C. and Elizabeth D. BoltonEdward Michael BonderBoston International TravelScott T. BradyLaura C. BroughtonSarah Dixwell Brown In memory of Edward Browning MeigsRobert H. Broyles In memory of Cornelius F. StrittmatterJudith A. Bruguiere In memory of Patricia A. Brown In honor of Christopher GentchosThomas and Jo Ann BylZoe and Andrew CardonMargaret W. and Samuel B. CarrMary CatanzaroRichard ChadwickChenbei ChangJohn A. ChristianVera S. ClarkLawrence B. CohenJoan and John Collier

Marc D. Coltrera and Anne L. BuchinskiHarry F. Conner and Carol E. Scott-ConnerNathaniel S. and Catherine E. CoolidgeJames Niles Coulombe In memory of Eric H. DavidsonNessly and Susan Craig In memory of John J. CebraMarilyn E. and J. Sterling CrandallCharles M. Crane and Wendy BreuerKaren CrawfordPrince S. Crowell, IIIJiamin CuiMary Ellen M. CunninghamRichard D. and Virginia M. CutlerRuth H. DavisNigel W. and Leila DawEllen P. DonaldsonLaneta and Mark DorflingerQuan-Yang Duh and Ann ComerDavid and Sherri DuricaLaurel A. EckhardtFrank Egloff, II In memory of Frank R. Lillie In honor of Frank EgloffAnn EhrenstromPaul T. Englund and Christine S. SchneyerDavid and Lois EpelDoris W. EpsteinMarnae Ergil In memory of Peter N. Kivy In honor of Joan PearlmanThomas E. EvansRobert R. and Ann EverettDonald** and Jo FaberKaren Fairbank-Moosekian In memory of Peter N. Kivy In honor of Joan PearlmanElizabeth M. FarnhamDavid and Doris L. FauschMarla B. FellerBarry and Shea Festoff In memory of Herbert S. FestoffAlan and Joan FinkelsteinMichael J. and Joann FishbeinFrederick S. and Ingeborg K. FisherHarvey M. and Susan Fishman

Karl W. Flessa and Mari N. JensenWendy FoxHadley Mack FrenchYoshiko Fujita In memory of Holger W. Jannasch and

Edward R. LeadbetterJohn J. FunkhouserHarold** and Ruth S. GainerDavid GallagherMichael R. and Mary GarfieldDouglas and Elizabeth GarlandC. Vernon GawAlan E. GellerStephen E. GellisAmy R. Genkins In memory of Peter N. Kivy In honor of Joan PearlmanSteven M. and Margaret R. Genkins In memory of Peter N. Kivy In honor of Joan PearlmanElizabeth L. GeorgeDuncan GibsonBarbara B. Glade In memory of Richard W. GladeJames A. GlazierMichael P. Goldring In memory of Irene GoldringJoel S. Gordon In memory of James W. LashMarion K. Gordon In memory of Robert B. Barlow and

David C. BeebeWerner M. GrafLewis J. GreeneDebra S. GregaHuson and Edith Gregory In honor of Paul J. De WeerLinda L. and Stephen A. GreyserAlbert and Sondra GrossmanZach W. Hall and Julia A. GiacobassiJohn F. and Sandra HalseyClifford V. Harding and Mina K. ChungGlenn W. HarringtonMary S. HarringtonKristen M. Harris and Max SnodderlyStephen C. Harrison

Jonathan and Dorothy Hasson In honor of Jeffrey PierceEdward E. HellerRobert and Taska HenerCatherine E. Hibbitt and Tim RockwellJohn** and Olivann HobbieRaymond W. HoltonJane HoltzRonald R. Hoy and Margaret C. NelsonHaruhiko ItagakiDieter JaegerErnest G. and Pauline B. JaworskiNancy A. JohnsonDaniel and Jean JohnstonElizabeth Ann Jonas and Thomas D. EisenFreda KaminerHung-Ying Kao and Gracie Hui-Chen

HuangMichael G. KauffmanRichard E. and Cynthia KendallThomas C. and Mary Ann KennyClifford H. Kern, IIILouie M. and Brenda KerrNasim KhanMarc W. and Phyllis KirschnerJenne KlotzJohn and Susan KrezoskiAndrew M. Kropinski and Peggy A.

PritchardAlan M. and B. J. KuzirianStory C.** and Dennis M. D. LandisMarnie and John LangfordMonica M. LarondaLouis S. and Camilla B. LarreySteven V. and Jessica G. LauriaBerton J. LeachMatthew K. LeeVanessa LeFevreDavid P. LenziEve and Ted LippoldPhilip and Martha LongVictoria H. and Francis C. Lowell In honor of Victoria LowellAnne LymanRobert E.** and Marjorie J. MainerRobert Paul Malchow


Phillip B. and Yvonne R. MaplesDawn E. MarickAndrew C. MarinucciSharyn MarksJunko Munakata MarrJulian B. MarshSandra K. Masur and Victor SchusterJane A. McLaughlinDavid W. McLayNawrie Meigs-Brown and David BrownEverett MendelsohnAllen F. MensingerWilhelm and Nonie MerckNancy S. and Richard Milburn In memory of Ruth Hubbard WaldRicardo* and Mela MilediDavid A. and Virginia I. MillerLynn Miller In memory of Edward R. LeadbetterRoberta MorrisHilary G. and Archie T. MorrisonDay O. and Kathie C. MountAngus Nairn and Marina PicciottoYasuko and Shigehiro NakajimaDavid E. NaylorPhil NehroPamela Nelson and Christopher Olmsted In memory of Leonard NelsonAnn Little NewburyPeter M. and Virginia NicholasSanto V. Nicosia In memory of Luigi MastroianniAlexander NikasAbigail NormanMichael and Carol Oberdorfer In memory of Robert B. BarlowJames L. and Nora OschmanRenee Bennett O’SullivanLorrence J. OtterArthur B. Pardee and Ann B. GoodmanJill J. Park In memory of James JeffersonJay and Nichole ParrishChee Pearlman In memory of Peter N. Kivy In honor of Joan Pearlman

Matthew A. Person and Jill EricksonBruce J. PetersonCarey R. and Ryan Petrie In memory of David L. GarbersGeorge H. and Louann S. PloughJeanne S. PoindexterCharles and Susan Porter In memory of Catherine N. NortonKenneth D. PossMatthew H. PowerDouglas PowersDonald and Helen PriceWilliam S. and Catherine A. ReznikoffFrederick R. and Kathryn G. RicklesPhilip S. and Anne RobertsonJames RobinsonDana and Alison RodinRichard E. RohrChristine K. Rolka In memory of William J. KuhnsEllen V. Rothenberg In memory of Eric H. DavidsonPriti Roy In honor of Prabhawati KumariMitch Rubin and Lisa BerkowerNorman B. RushforthMarie RussellMorton T. SaundersTina M. Saxowsky In memory of Elisabetta UlluThomas and Judith S. SbarraJay SchippersJoseph H. SchneiderThomas J. SchnitzerHerbert SchuelHarold H. and Kristin T. SearsHarriet F. SegalRobert and Caroline Seidler In memory of A. Kristine JohnsonTerrence J. SejnowskiSol Sepsenwol In memory of Teru HayashiGaius and Eleanore ShaverJerry W. ShayRichard and Elizabeth ShrinerAlan R. and Phyllis J. Silver

Roger D.** and Carol J. SlobodaJames SlockRoxanna S. and Ronald J. SmolowitzWilliam T. Speck and Evelyn LipperJohn W. SpeerLucy SteereAlan B. and Sala SteinbachElizabeth Stommel In memory of Henry T. StommelLaura C. StreichertKaren A. Summers In memory of William C. SummersSusan Szulman In memory of Aron E. SzulmanMargaret W. Taft In memory of Daniel S. GroschMarvin L. and Betsy C. TanzerPeter Tassia and Maija LutzCatherine D. ThalerWilliam R. ThomasMary and James ThorneNoah and Janet B. Totten In memory of Catherine N. NortonCatherine TrainorBarbara S. TrelaTsuchiura Daiichi High SchoolKenyon S. and Joan TweedellMichael and Frances W. TytellAmber Ulbrich In memory of H.B. SteinbachJoseph J. VallinoMiranda Van GelderKensal** and Barbara van HoldeRobert E. Van SciverCarol A. and Stephen P. WagnerRobin A. WallaceBrant G. WangJanis C. Weeks and William M. RobertsRobert H. and Marilyn WernerWayde M. WestonPaul S. and Barbara B. WheelerJustin and Jane WickensRoland L. and Nancy WigleyClayton A. and Arlene WileyBarbara Russell Williams In memory of Henry D. Russell

Carolyn Williams In memory of Howard B. LaneMarta G. WilliamsJames and Karen WycheCharles R. and Ellen G. WyttenbachWise and Lily YoungHarold H. Zakon and M. Lynne McAnellyRichard E. ZigmondJoshua J. Zimmerberg and Teresa L. Z. Jones

Up to $99Anonymous (2)Jaime R. Adame-GallegosSharon L. AmacherMaya Anjur-DietrichScott L. ApplebaumMike and Elana ArnoldMadelyn M. BaranMichael J. BarnesErnest BarretoOlive C. BeverlyDawn M. BlitzAshley Bolbrock In memory of Donna BochanowskiCharles and Helen BresnahanJennifer and David BrowerSusan M. Burden-GulleyLinda S. CarpenterRobert H. and Myra S. CarrierDonald L.D. CasparDanielle and Donald ChurillaJohn M. and Theresa M. ClarkinMort and Susan CohanChristopher and Karen CronanDorothy I. CrossleyMartha Bridge DencklaMichelle J. DoyleRobert A. DubinAllan and Linda DunnRita and Paul E. DussaultJudith S. EisenBrett Eric EtchebarneKristine FisherEdward H. FitchJohn W. FleegerJohn H. Ford


Peter FranklinNancy and Joseph V. FraserLarry Jay FriedmanFabrizio GabbianiAmy S. GladfelterSarah C. GoodrichAlison GreenJeffrey M. GrossLawrence I. and Esta S. GrossmanTom and Lisa Hammett In honor of Jeffrey PierceMichael and Joann Hanks In memory of Raymond and Rose EliottLyda S. HarrisRaymond and Elizabeth HayesMary G. HeardGina and Edmund A. HebertClare HeffernanLawrence and Layna Hobbie In honor of John E. HobbieKelly HoganJoan G. HoldenSeymour and Victoria E. HoltzmanBetsy and John HoneyLinda A. Hufnagel-Zackroff In memory of Ernest HufnagelMaryann HughesElaine M. HullCynthia B. HurleyCarolyn L. JohnsonClara KaoMelanie KarpM. Whitney KeenDoreen R. KellyDiana KenneyTilman J. KisperskyKathiann KowalskiBenjamin H. KrinskyMichael J. LandzbergDonna A. LaRocheSusan Laster and Terry LittleHerbert and Karen B. Levitan In memory of Steven LevitanChenghui LiEdward Locke In honor of Mother M. Locke

William John Robert LongabaughSusan LoucksAnne L. MacaulayMargaret M. MacLeish In memory of John CraneJane and Daniel MacNeilPatrick J. and Lorraine MahoneyCharles K. and Susanne J. MannDiane B. and Joseph S. MaranchieHannah MarkJoseph C. and Connie MartynaIsabelle McDonald In honor of Ann BowerWilliam and Kathleen McGoeyJoseph and Darlene McGuireCarol McKeonCornelia Hanna McMurtrieSamuel McMurtrie, Jr.Margaret A. Mills-Michael In memory of Harold M. MillsHazime MizoguchiYasu MoritaM. Patricia MorseThomas C. and Elizabeth D. MoseleyAnthony G. Moss In memory of Stewart and Dorothy MossKaren MrukStephen H. MunroeMary U. Musacchia and James E. FaberDenise Renee NelsonIris Nelson-Schwartz In memory of Lester and Lucena BarthBrian and Mary NickersonDavid OddeStephen H. OgilvyJeffery and Valerie OppenheimKathleen M. O’SullivanLeonard M. PassanoRonald J. and Dawn G. PfohlLouis Skjei PrahlChristian A. PutnamJeffrey L. RamShirley RapsLockwood RianhardMary Elizabeth RiceAlison A. Robb

Warren Rogers and Linda St. Jacques-Rogers

Mario RosascoRobert L. RosenbergRudi J. and Hanna RottenfusserLew* and Esther RowlandNathan C. RowlandE. Joel SafranMary L. ScaleraMary E. Schiffman In honor of Mother EmmonsSchool of the Holy ChildRalph T. SchwarzNeeraj SinghKathleen K. and Robert SiwickiPeter J.S. SmithCarmen S. SotoSven and Anna SpoerriJohn and Betsy StegemanAnn and Edward L. SternJudith G. StetsonFrank and Loretta TangJohn F. UrbanJoseph W. Vanable, Jr.Irving and Vida WagnerNancy WalbekYihong WangDeborah Morris WarnerAnthony Dale Weeks In memory of Gwen Szent-GyörgyiOlivia and Terry WhiteAshley V. WiresOliver H. WoshinskySusan WrayIsabel and Richard YoderMary Ruth YoeDorothy M. YorkAlexander and Judith S. Ziss

GIFTS TO THE MARINE BIOLOGICAL LABORATORY ALUMNI FUNDDonors to the Alumni Fund provide important support for the Marine Biological Laboratory’s Education Courses. In 2017, 533 graduate students from 39 countries and 342 institutions, 74 undergraduate students from 3 countries and 41 institutions, and 870 faculty, lecturers, and teaching assistants from 17 countries convened at the MBL.

Anonymous (6)C. Ronald AndersonRobert AngererErin and Barry AprisonElizabeth I. ArnoldMichael S. Ascher In honor of Shinya InoueRobert F. AshmanKatherine Bachman and Robert FlemingMichael Baltzley and Maia JonesRobert Barker and Marla Schay BakerNelson R. BartonEdward Joseph BehrmanJerry A. Bell and Mary Ann SteppGerald BergtromAri BerkowitzStanley J. BielenElizabeth A. Booth and Dustin L. DovalaAndrew BrittinghamBurt V. BronkStephen C. and Patricia S. BrownPatrick J. and Mary R. CalieAmy Camarillo In honor of David B. CamarilloGeorge A. ClarkAdam E. CohenAnnette W. and John R. ColemanStephen D. CollinsEdwin L. CooperJeffrey T. CorwinPamela A. CoutchieMr. John CunniffGregory Alan Dasch


Hannah A DeBergBruce and Judith Diner In honor of David and Miriam MauzerallThelma D. and Jonathan S. DixonDavid D. DowJudith S. EisenStanley Goodman ElfbaumAli ErogluArnold G. and Paula B. Eversole In memory of William D. Russell-HunterSuzanne Fachon-KalweitAlexandra S. FairfieldGeorge FeherKenneth Freedman and Steven WeissHugo D. and Anita R. FreudenthalMatthew FroschMatthew FrostWilliam Frost and Lise S. EliotAnne E. FryArthur G. GainesPaul E. GallantMaureen A. GannonSusan Gerbi-McIlwainHelen W. GjessingDavid L. GlanzmanDon Richard GoeMargaret Ann GoldsteinHolly V. GoodsonLeslie and James K. Gregg-JollyMatthew HackenbergKarl and E. Janet Hager In memory of Carolyn SlaymanSusan Harding HawkinsWilliam Allen HeldGordon E. HeringSarah E. Hitchcock-DeGregori In memory of George W. de VillafrancaRaymond W. HoltonTimothy E. Holy and Ieva M. SwansonEdward G. HornChristine Lee HoweRu-Chih C. HuangKachi Illoh In memory of Emmanuel and

Gertrude N. IllohJon W. Jacklet

Daniel E. JamesJohn P. JasperRena T. JonesErik M. JorgensenGordon I. KayeRobert G. KempR. Emmet and Gail KenneyLynda A. Kiefer In memory of Holger W. JannaschJennifer M. KingDavid Louis KirchmanAlan KlotzJames J. KnierimRobert E. KnowltonBirgit KovacsGeorge H. KowallisBruce B. LathamJanine M. LeBlanc-StraceskiWilliam J. LehmanArdean LeithEllen K. LeMosySteven William L’HernaultAnne M. LintonJennifer Lippincott-SchwartzHoward W. LowyEduardo R. and Nancy P. MacagnoEugene M. MartinWilliam H. MassoverThe MathWorksJoan MattsonAnne E. McBrideMegan N. McCleanMelanie and Klein MerrimanAnne Messer and Daniel GordonSara MichieLeonard E. MindichMartin M. Miner In memory of George T. ScottDavid M. and Theresa J. MiyamotoTetsuto MiyashitaPatrick L. Moore*Mary C. MullinsRuth H. NeffRalph F. NelsonPhyllis NorrisSara C. Pimental

Mark PlummerWilliam K. PlunkettWilliam F. Pomputius, IIIGeorge RadcliffeJane E. Radix In honor of Susan LeschineNancy RatnerJean F. and Ronald R. RegalVincent RehderRebecca F. ReuterMary Esther RicePierre E. Rouviere In memory of Carl WoesePriti Roy In memory of Veronica RodriguesUldis RozeKarl J. RusterholtzUeli RutishauserDavid R. SamolsRobert J. SaquetonAndrew SchutrumpfWenying ShouJames M. SidieKristina D. SimmonsPhilip J. Skehan, Jr.Alan M. SoohoAllan C. SpradlingJoel P. StafstromBarbara A. StayRobert E. SteeleJoan A. and Thomas A. SteitzCharles H. and Donna J. SullivanHarold E. SweetmanBen G. SzaroJosephine P.G. TanD. Lansing and M. Margaret TaylorSaul TeichbergCarl L. Thurman, IIJohn H. TietjenNatascia TisoLaurence C. TorsherAnn H. and Mark F. TraxAnthony VattaySusan Frances VolmanBrant G WangGary Ward and Zail Berry

Joseph T. WardenCory A. WatersWayde M. WestonHarold Bancroft WhiteCarmen J. Williams and Ann E. MillerHelene D. WinstanleyGeorge Wittenberg and Alyson H.

Bonavoglia In memory of Walter HeiligenbergRobert G. WolkKaren P YorkM. Jade Zee In memory of Allison DoupeSteven Zeichner


1888 SOCIETYThe 1888 Society recognizes and honors those who have made bequests and other deferred gift arrangements that will benefit the Marine Biological Laboratory in the future. We thank all of the members of the 1888 Society for their generosity.

Anonymous (7)Peter B.** and Margaret T. ArmstrongRobert B.** and Patricia Ann BarlowJoan R. BermanMargaret C.** and Francis P. BowlesZylpha B. BrownRonald Calabrese** and Christine

CozzensJeffrey D. CampbellGraciela C. Candelas*Adolph S. CavalloPaul N. ChervinJulia S. and Frank M.** ChildLawrence B. CohenSeymour S. CohenJohn E.** and Judith F. DowlingJoseph and Sarah DowlingPaul R. Dupee, Jr.**Barbara E. Ehrlich**David and Lois EpelMartha S. FergusonGerald D.** and Ruth L. FischbachBruce and Barbara FurieSusan Gerbi-McIlwainSol H. GoodgalPhilip Grant**Gladys GreenGertrude W. HinschJoseph F. HoffmanM Howard** and Frances F. JacobsonDaniel and Jean JohnstonSally S. and Ramsey E. JoslinJane Kaltenbach-Townsend*Darcy B. Kelley**Andrew M. Kropinski and Peggy A.

PritchardHans Laufer**

Paul B. Lazarow**Gloria A. LeadbetterMarian E. LeFevreJack** and Francine LevinHoward W. LowyThe Estate of Louise M. LuckenbillGordon MacIver and Eleanor MacIverRobert E.** and Marjorie J. MainerMiriam J. and David MauzerallMelanie and Klein MerrimanAnn E.** and John W.** MooreArthur B. Pardee and Ann B. GoodmanBarbara Standish PattenJeanne S. PoindexterThomas D.** and Patricia S. PollardRobert A. Prendergast**Ronald J. PrzybylskiMichael and Diane RabinowitzMarjorie ReinischPatricia** and Charles RobertsonHilde RosenthalMichael G. Ryan and Linda A. JoyceThe E. Joel Safran TrustSheldon J. ** * and Harriet F. SegalCecily Cannan SelbyEnid K. SichelPhilip J. Skehan, Jr.Roger D.** and Carol J. SlobodaMarjorie E. SteinbergJohn Tochko and Christina Myles-TochkoJohn J.* and Frederica W. ValoisSusan Frances VolmanEarl H. WeidnerJohn S. Willis

BEQUESTS REALIZED IN FY17Sybil Holtzer*Clare M. Wilbur*Leslie J.* and Elizabeth M.* Wilson

CORPORATE LENDING PROGRAMThe quality and success of the Marine Biological Laboratory’s educational program is maintained through research equipment, reagents, and computers valued at approximately $25 million loaned by:

89 North

Abberior Instruments AmericaAdvanced Microscopy GroupA-M SystemsACD Bio (Advanced Cell Diagnostics)AD InstrumentsAgilent TechnologiesALA Scientific Instruments, Inc.ALVEOLEAMETEK / TMCAndor/Bitplane/Oxford InstrumentsAngstrom Scientific / HitachiApplied Scientific InstrumentationAquaneeringATCCAutomate ScientificAxiom Optics

BD Life SciencesBeckman Coulter Life Sciences/Flow

cytometryBenedictus SystemsBio-Rad Laboratories, Inc. | Life Science

GroupBioptechs Inc.Bio-TechneBioVision TechnologiesBitplaneBoston Electronics CorporationBrain Vision LLCBruker

Cambridge Electronic Design LimitedCarl Zeiss Microscopy, LLCCharles River LaboratoriesCherry BiotechChroma

CoherentConoptics

Dagan CorpDiagenode / ThermoFisherDigitimerDoric LensesDPSS LasersDrummond Scientific

Electron Microscopy SciencesELITech Group / WescorEMD Millipore - CellasicEppendorf North AmericaEtaluma

Fine Science ToolsFluicellFluigentFormlabs

GE HealthcareGeneTools

HamamatsuHamilton ThorneHanna InstrumentsHarrick PlasmaHarvard BioscienceHewlet PackardHORIBA Scientific

iCub RobotImaging and Analysis, LLCImagine Optic Inc.Intelligent Imaging InnovationsInvitrogen

Jackson Laboratory

Keyence Corporation of AmericaKeysight Technologies

Laxco Inc.Leica MicrosystemsLI-COR Biosciences


Lonza WalkersvilleLuigs & NeumannLumencor, Inc.Luxendo / Bruker

MathWorksMatTek CorporationMBF BioscienceMeasurement ComputingMedical AssociatesMicrovideo Instruments Inc.MightexMiltenyi BiotecMolecular DevicesMolecular Probes

Nanoprobes, Inc.Narishige International USA, Inc.National InstrumentsNatus (formerly Grass Instruments)NeuralynxNeuroPhase LLCNew England BioGroupNew England BiolabsNewport CorporationNikon Instruments Inc.Noldus Information Technology

Optical Associates, Inc.OkoLab USA Inc.Omega Optical Inc.

Parker Hannifin CorporationPCO-TECH, Inc.Pearson Education - Math & SciencePentairPhi Optics, Inc.Photometrics / QimagingPrecisionary InstrumentsPrior ScientificPromega

Quest Scientific

Rapp OptoElectronic GmbHResearch Precision Instruments

Scientific Volume ImagingScientificaSeahorse BioscienceSemrockSiskiyou CorporationSolamere TGSpecialty Coating SystemsSpectra-Physics LasersSpectral EvolutionSPOT ImagingStoeltingSutter Instrument

Technical InstrumentsTED Pella, Inc.The MathWorks, Inc.Thermo FisherThermo Fisher Scientific / EVOS / Cellular

Imaging and Analysis Group | Life Science Solutions

ThorlabsTokai Hit Co.Toohey CompanyTucker-Davis Technologies

Unisense A/S

Viewpoint Behavior TechnologyVWR

Warner Instruments / Harvard Bioscience

Wavemetrics Inc. / IGOR ProWorld Precision Instruments

Yokogawa


LeadershipAs of June 30, 2017

TRUSTEE EMERITIFrederick Bay Norman Bernstein Margaret C. Bowles Seymour S. Cohen Mary B. Conrad Thomas S. Crane Paul R. Dupee Gerald D. Fischbach Prosser Gifford Robert D. Goldman Mary Greer Robert Haselkorn Ruth Hubbard* Kurt J. Isselbacher M Howard Jacobson George M. Langford Burton J. Lee* Irwin B. Levitan George W. Logan Robert E. Mainer Paul A. Marks Walter Massey Ambrose K. Monell R. Dana Ono Jean H. Pierce Thomas D. Pollard Robert A. Prendergast Frank Press Patricia K. Robertson Charles M. Rosenthal John W. Rowe Vincent J. Ryan James A. Sharp David Sheprowv Gerald Weissmann Christopher Minot Weld* Annette L. Williamson McColm Alfred M. Zeien

DIRECTOR TRUSTEE EMERITIGary G. Borisy John E. Burris Paul R. Gross Joan V. Ruderman

PRESIDENT TRUSTEE EMERITUSJohn E. Dowling

BOARD OF TRUSTEES Chairman Robert J. Zimmer, The University of Chicago

Vice Chair Saul J. Pannell, Wellington Management Company, LLC

President and Director Huntington F. Willard, Marine Biological Laboratory

Co-Interim Directors Melina E. Hale, The University of Chicago Neil Shubin, The University of Chicago

Treasurer Mary S. Harrington, Marine Biological Laboratory

Secretary Darren R. Reisberg, The University of Chicago

Chief Operating Officer Paul Speer, Marine Biological Laboratory

Trustees Robert H. Ament, Esquire, Ament Law Firm, LLC David B. Fithian, The University of Chicago Margaret J. Fofonoff, Art Lab LLC L. Patrick Gage, enGage Biotech Consulting James N. Galloway, The University of Virginia David A. Greene, Colby College H. David Hibbitt, Bristol, RI William I. Huyett, McKinsey & Co., Inc. Eric D. Isaacs, The University of Chicago Christopher G. Kennedy, Joseph P. Kennedy Enterprises Ethan A. Lerner, Massachusetts General Hospital Walter E. Massey, School of the Art Institute of Chicago John W. McCarter, Jr., The Field Museum Edward P. Owens, Norwell, MA Jeffrey H. Pierce, Pierce Aluminum Co., Inc. George Putnam, New Generation Advisors, LLC William T. Speck, New York, NY


Advisory Bodies

MBL COUNCILDonald R. Aukamp, Consulting Environmental EngineerDavid Bakalar, Physicist, Artist, and Film ProducerTimothy A. Barrows, Matrix Partners John F. Batter, III, WilmerHale George Berkowitz, Legal Sea Foods Marianne Billings, Friedman, Billings & Ramsey Ed Brakeman, Bain Capital, LP Joseph L. Dowling, The Rhode Island Eye Institute Jean-François Formela, Atlas Ventures Pamela L. Follett, Lewis Rhodes LabCraig B. Gibson, Lawrence General Hospital Gary Jacobson, Westwood Pembroke Health System Thomas P. Jalkut, Nutter, McClennen & Fish, LLPBarbara W. Jones, Barbara Woll Jones Designer/Builder Saul Kurlat, Kurlat Ventures Deena Gu Laties, ArtistGeorge V. Lauder, Harvard University Jennifer O. Letitia, IOMICS Pathways Initiative Joseph S. Levine, Concord, MAAnna Lysakowski, University of Illinois at Chicago William K. Mackey, Mackey & Foster, PA Dale Mathias, Africa Private Capital Group, USAIDRobert W. Pierce, Jr., Pierce Aluminum Co. John Ripple, Exonics Therapeutics David Schnell, Prospect Venture Partners Cecily C. Selby, CCS Ltd.Matthew L. Sherman, Acceleron PharmaJohn D. Sigel, WilmerHale Timothy A. Springer, Harvard Medical School Michael Stansky, OrthoSensor, Inc.Carolyn Stimpson, Wachusett Mountain Ann E. Sutherland, University of Virginia Gerard L. Swope, Woods Hole, MAJohn F. Swope, Concord, NHMary C. Swope, Woods Hole, MAKaren Tierney, Woods Hole, MAFrederica W. Valois, Woods Hole, MAJohn N. Vournakis,* Marine Polymer Technologies Nancy G. Wolf, Case Western University Barbara Wu, Wilmette, IL

PRESIDENT’S SCIENTIFIC ADVISORY COUNCILRobert Baughman, Okinawa Institute of Science and

Technology Graduate UniversityColleen Cavanaugh**, Harvard University, ex officioRita R. Colwell**, University of MarylandNancy Knowlton, Smithsonian InstitutionRobert (Robb) Krumlauf, Stowers Institute for Medical Research Jennifer Lippincott-Schwartz, Howard Hughes Medical Institute

– Janelia Research CampusMary C. Mullins, University of Pennsylvania Perelman School of

MedicineDavid C. Page, Whitehead InstituteDyann Wirth**, Harvard School of Public Health

MARINE BIOLOGICAL LABORATORY/THE UNIVERSITY OF CHICAGO AFFILIATION ADVISORY COMMITTEENeil Shubin, Department of Organismal Biology & Anatomy,

The University of Chicago (Chair)Conrad Gilliam, Dean for Research and Graduate Education,

Biological Sciences Division, The University of Chicago Jonathan Gitlin, Director, Division of Research, Marine

Biological LaboratoryMelina Hale, Department of Organismal Biology & Anatomy,

The University of ChicagoSusan Kidwell, Department of Geophysical Sciences, The University of ChicagoRae Nishi, Director, Division of Education; Burroughs Wellcome

Fund Director of EducationVicky Prince, Department of Organismal Biology & Anatomy;

Dean, Office of Graduate Affairs, The University of Chicago Matthew Tirrell, Director and Dean, Institute of Molecular

Engineering, The University of Chicago

SCIENCE COUNCIL OF THE MBL SOCIETY Tim Mitchison (Chair) Zoe Cardon Colleen Cavanaugh**, ex officioBill Green Melina Hale, ex officioRoger Hanlon Anne Hart

Len Kaczmarek Lee Niswander Ed Rastetter Neil Shubin, ex officioRoger Sloboda** Mitch Sogin Huntington F. Willard, ex officio


Scientists at MBL

RESIDENT SCIENTISTSLinda Amaral Zettler, Bay Paul CenterIrina Arkhipova, Bay Paul CenterZoe Cardon, Ecosystems CenterRichard Chappell, Bell CenterAnne Giblin, Ecosystems CenterJonathan Gitlin, Bell CenterKristin Gribble, Bay Paul CenterRoger Hanlon, Bell CenterMarko Horb, Bell CenterJulie Huber, Bay Paul CenterAlan Kuzirian, Bell CenterDavid Mark Welch, Bay Paul CenterJessica Mark Welch, Bay Paul CenterLydia Mäthger, Bell CenterJerry Melillo**, Ecosystems CenterJennifer Morgan, Bell CenterHilary Morrison, Bay Paul CenterRudolf Oldenbourg, Bell CenterEdward Rastetter, Ecosystems CenterLoretta Roberson, Bell CenterJoshua Rosenthal, Bell CenterGaius (Gus) Shaver, Ecosystems CenterMichael Shribak, Bell CenterMitchell Sogin, Bay Paul CenterJianwu (Jim) Tang, Ecosystems CenterTomomi Tani, Bell CenterMaki Tani, Bell CenterJoseph Vallino, Ecosystems Center

WHITMAN CENTER SCIENTISTS SUMMER 2016Veronica Acosta, University of the Incarnate WordSimon Alford, University of Illinois ChicagoSrdjan Antic, University of Connecticut Health CenterPeter Armstrong**, University of California, DavisGeorge Augustine, Nanyang Technological UniversityRobert Baker, New York University Medical CenterElaine Bearer, University of New MexicoMichael Bennett**, Albert Einstein College of MedicineMagdalena Bezanilla, University of Massachusetts, AmherstScott Brady, University of Illinois at Chicago College of MedicineDavid Burgess, Boston CollegeFred Chang, University of California, San FranciscoLawrence Cohen, Yale University

Sean Colin, Roger Williams UniversityDaniel Colon-Ramos, Yale UniversityJohn Costello, Providence CollegeScott Dawson, University of California, DavisJoseph DeGiorgis, Providence CollegeJohn Dubach, Massachusetts General HospitalKirsten Edepli, New York University Abu DhabiDavid Gadsby, The Rockefeller UniversityJesse Gatlin, University of WyomingDaniel Gibson, Worcester Polytechnic InstituteAndrew Gillis, University of CambridgeAntonio Giraldez, Yale School of MedicineAmy Gladfelter, University of North Carolina at Chapel HillRobert Goldman, Northwestern University, Feinberg School

of MedicinePhilip Grant**, National Institute of Neurological Disorders

and Stroke/National Institutes of HealthJessica Gray, Harvard Medical SchoolStephen Green, California Institute of TechnologyWilliam Green, University of ChicagoValentin Gribkoff, Yale University School of MedicineAvram Hershko, Technion- Faculty of MedicineMiguel Holmgren, National Institute of Neurological

Disorders and Stroke/National Institutes of HealthLiam Holt, New York UniversityKe Hu, Indiana UniversityAnthony Hyman, Max Planck Institute of Molecular Cell

Biology and GeneticsElizabeth Jonas, Yale UniversityErik Jorgensen, University of UtahLeonard Kaczmarek, Yale UniversityGen Kaneko, Yale UniversityU. Benjamin Kaupp, Center of Advanced European Studies

and Research Darcy Kelley**, Columbia UniversityKamran Khodakhah, Albert Einstein College of MedicineAlexey Khodjakov, Wadsworth CenterAkatsuki Kimura, National Institute of GeneticsJames Kittelberger, Gettysburg CollegePatrick La Rivière, University of ChicagoHans Laufer**, University of ConnecticutGilles Laurent, Max Planck Institute for Brain ResearchRodolfo Llinas**, NYU/Langone Medical CenterChristopher Lowe, Hopkins Marine Station of Stanford UniversityJane Maienschein, Arizona State UniversityJocelyn Malamy, University of Chicago


Robert Malchow, University of Illinois at ChicagoSatyajit Mayor, National Centre for Biological ScienceAllen Mensinger, University of Minnesota DuluthMark Miller, University of Puerto Rico Medical

Sciences CampusAndrew Miller, Hong Kong University of Science

and TechnologyPaul Minor, Hopkins Marine Station of Stanford UniversityBrian Mitchell, Northwestern UniversityTimothy Mitchison, Harvard Medical SchoolGerardo Morfini, University of IllinoisJohn Oakey, University of WyomingHarish Pant, National Institute of Neurological

Disorders and Stroke/National Institutes of HealthEduardo Perozo, University of ChicagoSabine Petry, Princeton UniversitySimone Reber, Humboldt University of BerlinThomas Reese**, National Institute of Neurological

Disorders and Stroke/National Institutes of HealthAntonina Roll-Mecak, National Institute of

Neurological Disorders and Stroke/National Institutes of Health

Lawrence Rome, University of PennsylvaniaMichael Rosen, UT Southwestern Medical CenterWilliam Ross, New York Medical CollegeHari Shroff, National Institutes of HealthRoger Sloboda**, Dartmouth CollegeGreenfield Sluder, University of Massachusetts

Medical SchoolTimothy Springer, Boston Children’s HospitalThomas Surrey, The Francis Crick InstituteMark Terasaki, University of Connecticut Health CenterRon Vale, University of California, San FranciscoDaniel Ward, University of ConnecticutClare Waterman, National Heart, Lung, and Blood

Institute/National Institutes of HealthAllan Wiechmann, University of Oklahoma Health

Sciences CenterBrandon Weissbourd, California Institute of TechnologyJim Wilhelm, University of California, San DiegoJoshua Zimmerberg, National Institutes of HealthSteven Zottoli, Williams College

MBL FELLOWSDaniel Colon-Ramos, Yale UniversityMaureen Conte, Bermuda Institute of Ocean Science

Linda Deegan, Woods Hole Research CenterA. Murat Eren, University of ChicagoJack Gilbert, University of ChicagoAmy Gladfelter, University of North Carolina at Chapel HillMelina Hale, University of ChicagoRobert Howarth, Cornell UniversityPatrick La Rivière, University of Chicago

Jane Maienschein, Arizona State UniversityChristopher Neill, Woods Hole Research CenterClifton Ragsdale, University of ChicagoAlejandro Sanchez Alvarado Stowers, Institute;

Howard Hughes Medical InstituteHari Shroff, National Institutes of Health

GRASS FELLOWS SUMMER 2016Arne Battefeld, Netherlands Institute for NeuroscienceAriadna Cobo-Cuan, Havana UniversityChristophe Dupre, Columbia UniversityHelen Hou, Harvard Medical SchoolWanhe Li, The Rockefeller UniversityJustin O’Hare, Duke UniversityOlga Penagarikano, University of the Basque Country,

School of MedicineMatthias Prigge, Weizmann Institute of ScienceAlma Rodenas-Ruano, Fordham UniversityJoshua Salvi, The Rockefeller UniversityNan Wang, Gent University

COURSE DIRECTORS AND FACULTY SUMMER 2016Kjersti Aagaard, Baylor College of MedicineLarry Abbott, Columbia UniversitySara Abrahamsson, The Rockefeller UniversityJan Ache, Howard Hughes Medical Institute - Janelia

Research CampusCarlos Aizenman, Brown UniversityJuan Alarcon, SUNY Downstate Medical CenterDirk Albrecht, Worcester Polytechnic InstituteMark Alkema, University of Massachusetts Medical SchoolKelsey Allen, Massachusetts Intitute of TechnologyAndrew Alspaugh, Duke University School of MedicineSharon Amacher, The Ohio State UniversityJames Angstadt, Siena College

Ricardo Araneda, University of MarylandDetlev Arendt, European Molecular Biology LaboratoryMario Ascoli, The University of IowaStephen Baccus, Stanford UniversityShernaz Bamji, University of British ColumbiaAndrei Barbu, Massachusetts Institute of TechnologyDiana Bautista, Diana BautistaChristine Beattie, Ohio State UniversityPeter Beerli, Florida State UniversityRichard Behringer, University of Texas MD Anderson

Cancer CenterNathan Belliveau, California Institute of TechnologyJoanne Berger-Sweeney, Trinity CollegeDaniel Bernard, McGill UniversityJoseph Bielawski, Dalhousie UniversityDawn Blitz, Miami UniversityJames Boedicker, University of Southern CaliforniaJustin Bois, California Institute of TechnologyPawel Borowicz, North Dakota State UniversityHans Brecht, Humboldt University BerlinKris Brewer, Massachusetts Institute of TechnologyCarlos Brody, Princeton UniversityEmery Brown, Massachusetts Institute of TechnologyJoann Buchanan, Allen Institute for Brain ScienceDirk Bucher, New Jersey Institute of TechnologyRonald Calabrese**, Emory UniversityDavid Carroll, Florida Institute of TechnologyEdward Castaneda, The University of Texas at El PasoFlaminia Catteruccia, Harvard T.H.Chan School of

Public HealthMaurice Chacron, McGill UniversityDmitri Chklovskii, Simons FoundationLane Christenson, Lane ChristensonSueyeon Chung, Harvard UniversityJose Cibelli, Michigan State UniversityPaula Cohen, Cornell UniversityAndres Collazo, California Institute of TechnologyDaniel Colon-Ramos, Yale UniversityAlan Conley, University of California, DavisRobert Cramer, Geisel School of Medicine at DartmouthSean Crosson, University of ChicagoGraeme Davis University of California, San FranciscoScott Dawson, University of California, DavisBernard Degnan University of QueenslandChris Deister Brown UniversityMauricio Delgado, Rutgers University


Francesco Demayo, National Institute of Environmental Health Sciences

Sophie Deneve, Simons InstituteElva Diaz, University of California, Davis School

of MedicineJames Dicarlo, Massachusetts Institute of TechnologyGeorge Dimopoulos, Johns Hopkins UniversityIna Dobrinski, University of CalgaryPaul Dudchenko, University of StirlingOlga Dudko, University of California, San DiegoFrancesca Duncan, University of Kansas Medical CenterCasey Dunn, Brown UniversityBenjamin Eaton, The University of Texas Health

Science Center at San AntonioUri Eden, Boston UniversityRobert Edgar Scott Edwards, Harvard UniversityJohn Edwards, Harbor-UCLA Medical CenterEmmeline Edwards, National Institutes of HealthBuffy Ellsworth, Southern Illinois UniversityA. Murat Eren, University of ChicagoBard Ermentrout, University of PittsburghAnne Etgen, Albert Einstein College of MedicineJames Fadool, Florida State universityAdrienne Fairhall, University of WashingtonSteven Farber, Carnegie InstitutionAsgi Fazleabas, Michigan State UniversityMichale Fee, Massachusetts Institute of TechnologyAndre Fenton, New York UniversityAretha Fiebig, University of ChicagoIla Fiete, The University of Texas at AustinRobert Fischer National Heart, Lung, and Blood InstituteEva Fischer, Harvard UniversityRafael Fissore, University of MassachusettsJames Fitzgerald, Harvard UniversityFrancisco Flores, Massachusetts Institute of TechnologyEric Fortune, New Jersey Institute of TechnologyLoren Frank, University of California, San FranciscoWinrich Freiwald, The Rockefeller UniversityKathleen French, University of California, San DiegoFriedrich Frischknecht, Heidelberg UniversityGreg Gage, Backyard BrainsBrigitte Galliot University of GenevaSurya Ganguli, Stanford UniversityHernan Garcia, University of California, BerkeleyEthan Garner, Harvard University

Maria Geffen, University of PennsylvaniaSamuel Gershman, Harvard UniversityMatthew Gibson, Stowers Institute for Medical ResearchAmy Gladfelter, University of North Carolina at Chapel HillMark Goldman, University of California, DavisBob Goldstein, University of North Carolina at Chapel HillJacqueline Gottlieb, Columbia UniversityMichael Granato, University of PennsylvaniaStephan Grill, Technical University DresdenEyal Gruntman, Howard Hughes Medical InstituteHugo Guerrero-Cazare, Johns Hopkins UniversityGreg Hale, Massachusetts Insitute of TechnologyMarc Hammarlund, Yale School of MedicineStephen Hammes, University of RochesterKurt Hanselmann, Swiss Federal Institute of TechnologyRichard Harland, University of California, BerkeleyRayna Harris, The University of Texas at AustinJosh Hawk, Yale UniversityJames Haxby, Dartmouth CollegeTracy Heath, Iowa State UniversityJoseph Heitman, Duke University Medical CenterJonathan Henry, University of IllinoisMichael Higley, Yale UniversityJohn Hildebrand, University of ArizonaEllen Hildreth, Wellesley CollegeDavid Hillis, University of TexasBarry Hinton, University of VirginiaHans Hofmann, The University of Texas at AustinDeborah Hogan, Geisel School of Medicine,

Dartmouth CollegeMark Holder, University of KansasScott Holley, Yale UniversitySusan Holmes, Stanford UniversityMichael Hoppa, Dartmouth CollegeDavid Horn, University of DundeeCorinne Houart, King’s College LondonGreg Huber, University of CaliforniaJohn Huelsenbeck, University of California, BerkeleySusan Huse, Indigo AgricultureChristopher Huston, University of VermontCurtis Huttenhower, Harvard Chan SchoolLeyla Isik, Massachusetts Institute of TechnologyLaurinda Jaffe, University of Connecticut Health CenterRoshan Jain, Haverford CollegeGregory Johnson, Texas A&M UniversityKevin Jones, Howard University

Erik Jorgensen, University of UtahJoan Jorgensen, University of WisconsinPatrick Kanold, University of MarylandHsin-yi Kao, New York UniversityRolf Karlstrom, University of MassachusettsBoris Katz, Massachusetts Institute of TechnologyRaymond Keller, University of VirginiaJiye Kim, Boston Childrens HospitalJean King, University of Massachusetts Medical SchoolNicole King, University of California, BerkeleyRob Knight, University of California, San DiegoJane Kondev Brandeis UniversityJames Konopka, Stony Brook UniversityNancy Kopell, Boston UniversityGabriel Kreiman, Harvard Medical SchoolWilliam Kristan, University of California, San DiegoDamian Krysan, University of Rochester Medical CenterLaura Kubatko, The Ohio State UniversityMorgan Langille, Dalhousie UniversityRosamund Langston, University of DundeeJared Leadbetter, California Institute of TechnologyAlba Ledesma, National Institute of

Aropecuarian TechnologyDaniel Lee, University of PennsylvaniaHeun Lee, California Institute of TechnologyPaul Lewis, University of ConnecticutXiaorong Lin, Texas A&M UniversityTheodore Lindsay, California Institute of TechnologyJennifer Lippincott-Schwartz, Howard Hughes Medical

Institute - Janelia Farm Research CampusMary Logan, Oregon Health and Science UniversityPng Loke, NYU School of MedicineChristopher Lowe, Stanford UniversityEllen Lumpkin, Columbia UniversityPaul Maddox, University of North Carolina, Chapel HillL Mahadevan, Harvard UniversityEve Marder, Brandeis UniversityAndres Maricq, University of UtahWallace Marshall, University of California, San FranciscoMark Martindale, University of FloridaJoe Martinez, University of IllinoisDavid Matus, Stony Brook UniversityDavid McClay, Duke UniversityJosh McDermott, Massachusetts Institute of TechnologyEmily Jane McTavish, University of KansasLisa Mehlmann, UConn Health


Ethan Meyers, Hampshire CollegeKenneth Miller, Columbia UniversityThomas Misgeld, Technical University of MunichAaron Mitchell, Carnegie Mellon UniversityWiktor Mlynarski, Massachusetts Institute of TechnologyCecilia Moens, Fred Hutchinson Cancer Research CenterChristopher Moore, Brown UniversityTirin Moore, Stanford UniversityMartin Mueller, University of ZurichChristian Mueller, Simons FoundationFerenc Mueller, University of BirminghamLouis Muglia, Cincinnati Children’s Hospital Medical CenterMary Mullins, University of PennsylvaniaFarzan Nadim, New Jersey Institute of TechnologyDianne Newman, California Institute of TechnologyBrian Norris, Emory UniversityJodi Nunnari, University of California, DavisElke Ober, University of CopenhagenAude Oliva, Massachusetts Institute of TechnologyGeorge Otoole, Geisel School of MedicineCarolyn Ott, Howard Hughes Medical Institute -

Janelia Research CampusJay Parrish, University of WashingtonWilliam Pate, Ad Hoc AnalyticsNipam Patel, University of California, BerkeleyWilliam Pearson, University of VirginiaDarcy Peterka, Columbia UniversityMargaret Petroff, Michigan State UniversityRobert Phillips, California Institute of TechnologyTatjana Piotrowski, Stowers Institute for Medical ResearchGina Poe, University of MichiganTomaso Poggio, Massachusetts Institute of TechnologyKatherine Pollard, Gladstone InstitutesElisa Pompeo, Massachusetts Institute of TechnologyMihai Pop, University of MarylandKenneth Poss, Duke University School of MedicineKara Pratt, University of WyomingJoshua Puhl, University of MinnesotaChristopher Quince, University of WarwickAlfredo Quinones-Hinojosa, Johns Hopkins UniversityGregory Quirk, University of Puerto Rico School

of MedicineDavid Raible, University of WashingtonErez Raz, University of MuensterManuel Razo-Mejia, California Institute of TechnologyHesper Rego, Harvard University

Michael Reiser, Howard Hughes Medical InstituteLawrence Reynolds, North Dakota State UniversityGemma Roig, Massachusetts Institute of TechnologyMatthew Ronshaugen, University of ManchesterLorenzo Rosasco, Massachusetts Institute of TechnologyVanessa Ruta, The Rockefeller UniversityTimothy Ryan, Weill Cornell Medical CollegeJon Sack, University of CaliforniaAlejandro Sanchez Alvarado, Stowers Institute for

Medical ResearchStephan Sanders, University of California, San FranciscoMandana Sassanfar, Massachusetts Institute

of TechnologyAlex Schier, Harvard UniversityDavid Schulz, University of Missouri-ColumbiaElaine Seaver, University of FloridaTerrence Sejnowski, Salk InstituteDonald Sheppard, McGill University Health CenterDavid Sherwood, Duke UniversityHari Shroff, National Institutes of HealthNeil Shubin, University of ChicagoPhotini Sinnis, Johns Hopkins School of Public HealthSara Solla, Northwestern UniversityLilianna Solnica-Krezel, Washington University School

of MedicineHaim Sompolinsky, The Hebrew University of JerusalemSean Speese, OHSU-Jungers Center for

Neurosciences ResearchElizabeth Spelke, Harvard UniversityThomas Spencer, University of MissouriAngelike Stathopoulos, California Institute of TechnologyRob Steele, University of California, IrvineWolfgang Stein, Illinois State UniversityClaudio Stern, University College LondonAndrea Streit, King’s College LondonMichael Stryker, University of California, San FranciscoDavid Swofford, Duke UniversityLidia Szczupak, Universidad de Buenos AiresTracy Teal, Data CarpentryJamie Theobald, Florida International UniversityJulie Theriot, Stanford University School of MedicineManuel Thery, Hopital Saint-LouisKrista Todd, University of California, San DiegoJames Townsel, Meharry Medical CollegePaul Trainor, Stowers Institute for Medical ResearchKeith Trujillo, California State University San Marcos

Matthijs Van Der Meer, Dartmouth CollegeGiel Van Dooren, Australian National UniversityWesley Van Voorhis, University of WashingtonJohn Wallingford, University of Texas, AustinGary Ward, University of VermontShigeki Watanabe, Johns Hopkins UniversityDavid Weisblat, University of CaliforniaMelanie Wellington, University of Rochester Medical CenterTheodore White, University of Missouri – Kansas CityAthula Wikramanayake, University of MiamiCarmen Williams, National Institute of Environmental

Health SciencesEmma Wilson, University of California, RiversideDaniel Wolpert, University of CambridgeEmma Wood, University of EdinburghTerrence Wright, California State University, SacramentoDavid Wu, University of ChicagoRyohei Yasuda, Max Planck Florida Institute

for NeuroscienceDeborah Yelon, University of California, San DiegoAnne Yoder, Duke UniversityRafael Yuste, Columbia UniversityRobert Zeller, San Diego State UniversityYi Zuo, University of California, Santa CruzDerrick Zwickl, University of ArizonaDaniel Zysman, Massachusetts Institute of Technology

*deceased**former Trustee


PhotosCover: Pygmy Squid Idiosepius paradoxus (Tom Kleindinst)Inside front cover: MBL campus from the Belltower Gardens

(Emily Zollo)P1: Interim Co-Directors Melina Hale and Neil ShubinP2: L to R: David Mark Welch (Emily Zollo); Jennifer

Morgan (Emily Zollo); California Two-Spot Octopus Octopus bimaculoides (Tom Kleindinst); MBL Distinguished Scientists: John Dowling, Ronald Vale, Joan Ruderman, Avram Hershko, Shinya Inoué, Mitchell Sogin (Elizabeth Armstrong)

P3: L to R: Toolik Lake LTER (Jim Laundre); Caroline Albertin; Kristen Hunter-Cevera; Duygu Özpolat (Aaron Pomerantz); Physical Biology of the Cell student Lokesh Pimpale and course co-drector, Hernan Garcia (Megan Costello)

P4: L to R: Ronald Vale (Ryan Anson / AP Images for HHMI); Marko Horb, Marcin Wlizla, Sean McNamara (Emily Zollo); Color-coded actin filament orientation overlaid on an image of fluorescence intensity (Shalin Mehta and Tomomi Tani)

P5: Little Sippewissett Marsh (Megan Costello)

P7: UChicago student Jennifer Teng (Megan Costello)P8: Jeffery Pierce (Dan Cutrona)P21: Aerial view of Woods Hole and MBL campus (Matt Barton)Inside back cover: Fluorescent light micrograph of two

mouse fibroblast cells. (Torsten Wittmann)

2016 -2017 MBL ANNUAL REPORT i i i

MARINE BIOLOGICAL LABORATORY 7 MBL StreetWoods Hole, MA 02543www.mbl.edu

2016-2017 ANNUAL REPORT - Marine Biological Laboratory

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Transcript of 2016-2017 ANNUAL REPORT - Marine Biological Laboratory