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Cretaceous Research 30 (2009) 496–502

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Cretaceous Research

journal homepage: www.elsevier .com/locate/CretRes

Report on the 3rd International Meeting of the IUGS Lower CretaceousAmmonite Working Group, the ‘‘Kilian Group’’(Vienna, Austria, 15th April 2008)

Stephane Reboulet a,* and Jaap Kleinb (reporters) and Ricardo Barragan c, Miguel Company d,Celestina Gonzalez-Arreola c, Alexander Lukeneder e, Seyed Naser Raisossadat f, Jose Sandoval d,Ottilia Szives g, Jose M. Tavera d, Zdene�k Vasıcek h, Jean Vermeulen i

a UFR Sciences de la Terre, UMR-CNRS 5125 Paleobiosphere-Paleoenvironnements, Universite de Lyon, UCB Lyon 1, Batiment Geode, rue Raphael Dubois,69622 Villeurbanne cedex, Franceb Demmerik 12, 3645 EC Vinkeveen, The Netherlandsc Departamento de Paleontologia, Instituto de Geologıa, UNAM, Ciudad Universitaria, Coyoacan. Codigo Postal 04510, Mexico, D.F., Mexicod Departamento de Estratigrafia y Paleontologia, Facultad de Ciencias, Universidad de Granada, Campus de Fuentenueva s/n, 18002 Granada, Spaine Geological-Palaeontological department, Natural History Museum, Burgring 7, A-1010 Vienna, Austriaf Geology Department, Faculty of Science, Birjand University, PO Box 97175/615, Birjand, Irang Hungarian Natural History Museum, Department of Paleontology, 1088 Ludovika ter 2, H-1431 Budapest, Hungaryh Institute of Geonics, Academy of Sciences of the Czech Republic, Studentska 1768, CZ-70800 Ostrava-Poruba, Czech Republici Grand Rue, 04330 Barreme, France

a r t i c l e i n f o

Article history:Received 16 October 2008Accepted in revised form 12 December 2008Available online 24 December 2008

Keywords:AmmonitesZonationHauterivianBarremian

* Corresponding author.E-mail address: stephane.reboulet@univ-lyon1.fr (

0195-6671/$ – see front matter � 2008 Elsevier Ltd.doi:10.1016/j.cretres.2008.12.009

a b s t r a c t

The discussion is mainly focused on the Hauterivian and Barremian zonal schemes. For the HauterivianStage, the main changes concern the Subsaynella sayni and Balearites balearis Zones with the introductionof one horizon (Subsaynella begudensis Horizon) and four subzones (B. balearis, Binelliceras binelli, Crio-ceratites krenkeli, Spathicrioceras seitzi subzones), respectively. For the Lower Barremian, the mainamendments concern the introduction of two subzones in the Taveraidiscus hugii auctorum Zone (T. hugiiauctorum and Psilotissotia colombiana subzones) and of four horizons in the Kotetishvilia compressissimaZone (Holcodiscus fallax, Nicklesia didayana, Heinzia communis and Subtorcapella defayae Horizons). TheCoronites darsi Zone is replaced by the Moutoniceras moutonianum Zone in which two horizons occur (C.darsi and Heinzia caicedi Horizons). The base of the Upper Barremian is defined by the base of theToxancyloceras vandenheckii Zone and Subzone. The former Hemihoplites feraudianus and Martelitessarasini Zones are lowered in rank to subzones of the Gerhardtia sartousiana and Imerites giraudi Zones,respectively.

� 2008 Elsevier Ltd. All rights reserved.

1. Introduction

The workshop of the IUGS Lower Cretaceous AmmoniteWorking Group was planned the day before the first InternationalMeeting on Correlation of Cretaceous Micro-and Macrofossils(16–18 April 2008), both held at the Natural History MuseumVienna. Sincere thanks are extended to Alexander Lukeneder forthe local organization of the Kilian Group meeting. The workshopwas attended by 12 members from 8 countries.

Before starting discussion on the Standard zonation, thechairman made some comments and recommendations on the

S. Reboulet).

All rights reserved.

working method. Firstly, new proposals made by members of theKilian Group should be preferentially based on published orsubmitted data which have been reviewed or discussed previ-ously by some members of the Kilian Group (for example see the2007 Digne report, Reboulet et al., 2007). New data presentedduring a meeting do not allow to have enough time for reflection.Secondly, it would be preferable to keep zones which define thebase of stages or substages as they have been recommended bythe First (Birkelund et al., 1984) and Second (Mutterlose et al.,1996; Rawson et al., 1996 for Hauterivian and Barremian Stages,respectively) International Symposiums on Cretaceous StageBoundaries or chosen in the 2004 Geological Time Scale (Ogget al., 2004). Thirdly, it appeared necessary to stabilize the Hau-terivian-Barremian zonal scheme which has been often modified

S. Reboulet and J. Klein (reporters) et al. / Cretaceous Research 30 (2009) 496–502 497

during the last 8 years (Hoedemaeker and Rawson, 2000;Hoedemaeker et al., 2003; Reboulet et al., 2006; this report).Fourthly, members of the Kilian Group should use the Standardzonation or correlate it with their local zonal scheme. Thesedifferent points will allow to preserve the stability in the strati-graphic nomenclature (particularly for the first zones of sub/stages), to make easier the comparison of zonal schemes betweendifferent works and to improve communication. The aim of theKilian Group is to define a Standard zonation but also toencourage other colleagues to use it. Consequently, editors ofjournals will be informed by the chairman that authors shouldused the Lower Cretaceous Ammonite Standard zonation (when itis possible) or to add it beside the local zonal scheme.

The discussion mainly focused on the Hauterivian and Barre-mian zonal schemes. For each zone, Stephane Reboulet presenteda brief summary of amendments made by the successive LowerCretaceous Cephalopod Teams (Hoedemaeker et al., 1990, 1993,1995; Rawson et al., 1999; Hoedemaeker and Rawson, 2000) andthe Kilian Groups (Hoedemaeker et al., 2003; Reboulet et al.,2006) in order to put forward some important points or problems(this document, distributed at the meeting, can be asked to theauthor/chairman). The main results of the 2007 Digne meeting(Reboulet et al., 2007) were evoked and the comments of PhilipHoedemaeker and Pierre Ropolo about this report were alsogiven. Some proposals were made in order to give a base ofdiscussion. Several amendments were introduced and changes areshown in Tables 1 and 2; the numbers in the text refer to thenumbers in these Tables.

2. The standard zonation

2.1. Hauterivian

1. Acanthodiscus radiatus Zone. No change. As the onlycomment on this zone concerned the introduction of a Breis-trofferella castellanensis Horizon (Hoedemaeker and Rawson,2000), it is suggested to give further information taking intoaccount that the base of A. radiatus Zone is used to define thebase of the Hauterivian Stage. The La Charce section (Drome,France) is the proposed GSSP (Global Standard Stratotype Sectionand Point) for the Hauterivian Stage (Ogg et al., 2004; Rawson,2004). The Golden Spike is not yet assigned, but would probablybe placed at the base of layer 189 with the lowest occurrence ofAcanthodiscus rebouli (¼ first appearance of the genus, Reboulet,1996); this definition was recommended by the 1995 Brusselsmeeting (Mutterlose et al., 1996) and reported in the 2004Geologic Time Scale (Ogg et al., 2004).

2. Crioceratites loryi Zone. No change. During the 2000 Viennameeting (Hoedemaeker and Rawson, 2000), the former C. loryi andOlcostephanus (Jeannoticeras) jeannoti Horizons (Hoedemaekeret al., 1993) were elevated to subzones, but no explanation andreference were given about the palaeobiogeographic and strati-graphic distributions of these index-species. C. loryi and O. (J.)jeannoti have been recognized in numerous countries (Klein, 2005;Klein et al., 2007) and have a large potential of correlation. They arerelatively frequent or abundant in deep-water facies (Vocontianbasin) and seem less abundant in shallow-water palaeoenviron-ments (Provence and Jura platforms, Bulot, 1995; Reboulet, 1996and references).

3. Lyticoceras nodosoplicatum Zone. No change. For the Olcoste-phanus (Olcostephanus) variegatus Horizon, it was noted that thefirst appearance of the index-species is a few beds below the firstappearance of L. nodosoplicatum and thus the base of the horizonshould be drawn a little below the base of the L. nodosoplicatumZone (Hoedemaeker et al., 2003).

4. Subsaynella sayni Zone. The Subsaynella begudensis Horizonproposed by Vermeulen is adopted. Its base should be defined bythe first appearance of the index-species and not by the base ofthe acme (epibole) as proposed by Vermeulen et al. (1999). Thishorizon fills the gap between the top of the Cruasiceras cruasenseHorizon and the base of the Plesiospitidiscus ligatus Zone.C. cruasense and S. begudensis have been found only in SoutheastFrance (Vermeulen et al., 1999; Klein, 2005; Vermeulen and Klein,2006).

5. Plesiospitidiscus ligatus Zone. During the Mula meeting (Spain,1992), the inverted commas were added to mean that the index-species was not a true P. ligatus d’Orbigny, but P. ligatus sensuMoullade and Thieuloy, 1967 (Hoedemaeker et al., 1993). They havebeen deleted without explanation in the 2000 Vienna report(Hoedemaeker and Rawson, 2000). The concept of the index-species is now well known. The holotype has been figured(Vermeulen et al., 1999; Busnardo et al., 2003; Busnardo in Fischerand Gauthier, 2006) and the extension and content of the zone hasbeen correctly established (Vermeulen, 2003). P. ligatus sensuMoullade and Thieuloy is probably Plesiospitidiscus subdifficilewhich appears later than P. ligatus (Vermeulen, 2003; see alsoVermeulen in Reboulet et al., 2007).

6. Balearites balearis Zone. It is divided into B. balearis, Binelli-ceras binelli, Crioceratites krenkeli and Spathicrioceras seitziSubzones (using classification of Klein et al., 2007; according toCompany (in Klein et al., 2007), Spathicrioceras is hardly separablefrom Crioceratites). This intra-zonal scheme is mainly based on theproposal of Company et al. (2002, 2003) who suggested to subdi-vide this zone in 4 subzones defined by the consecutive appear-ances of 4 different chronospecies belonging to the sameCrioceratites lineage: C. balearis, C. binelli, C. krenkeli and C. anguli-costatus. Jean Vermeulen recognizes the first three subzones.However, he considers (see also Vermeulen in Reboulet et al., 2007)that C. angulicostatus sensu Company et al. (2003) is a S. seitzi(Sarkar, 1955). The S. seitzi and C. angulicostatus Subzones repre-sents the same interval. Indeed, Vermeulen (2003) correlated thebase of his former Spathicrioceras angulicostatum Zone with thebase of the C. angulicostatus Subzone, and more recently Vermeulen(2004) considered that specimens identified as S. angulicostatum(Kilian, 1889) are probably S. seitzi (Sarkar, 1955). After discussion,Miguel Company and Jean Vermeulen agree to use S. seitzi as index-species to name the fourth subzone.

B. balearis, B. binelli, C. krenkeli have a large distribution (Kleinet al., 2007). On the Provence platform (Autran, 1993), B. binelli andC. krenkeli occur only at the uppermost part of the B. balearis Zonewhich is characterized by the range of the index-species. In deeper-waters palaeoenvironments (Veveyse de Chatel area, Switzerland;Vocontian basin, France), the B. balearis-B. binelli succession wasalso observed (Busnardo et al., 2003; Busnardo in Reboulet et al.,2007). S. seitzi has been found in Switzerland and France (Kleinet al., 2007).

7. ‘‘Pseudothurmannia ohmi’’ Zone. It is recommended to addinverted commas to mean that P. ohmi awaits a redefinition. The P.ohmi Zone and the P. ohmi, Pseudothurmannia catulloi and Pseudo-thurmannia picteti Subzones are kept as proposed by Hoedemaekeret al. (2003) and Reboulet et al. (2006). Several discussions(Reboulet et al., 2007; Hoedemaeker’s comments; this workshop)have shown that further investigations are necessary in order toprecise the conception, identification and stratigraphic ranges ofthese index-species and other species as Pseudothurmanniamortilleti. Different interpretations would be also partly due tocollection failures. To reach an agreement, the chairman proposedto organize a workshop (probably in France, 2009), which will berestricted to colleagues who work on the uppermost Hauterivianammonite fauna.

Table 1Ammonite Standard zonation of the Berriasian-Hauterivian Stages. The numbers in this Table refer to the numbers in the text.

STAGES ZONES SUBZONES HORIZONSPseudothurmannia picteti

Pseudothurmannia catulloi "Pseudothurmannia ohmi" (7)"P. ohmi" (7)

Spathicrioceras seitzi (6)Crioceratites krenkeli (6)

Binelliceras binelli (6)Balearites balearis

B. balearis (6)Plesiospitidiscus ligatus (5)

Subsaynella begudensis (4)

Upper

Subsaynella sayniCruasiceras cruasense

Lyticoceras nodosoplicatum Olcostephanus (Olcostephanus)variegatus (3)

Olcostephanus (Jeannoticeras)jeannoti (2)Crioceratites loryi C. loryi (2)

HA

UT

ER

IVIA

N

Lower

Acanthodicus radiatus (1) Breistrofferella castellanensis Teschenites callidiscus

Criosarasinella furcillataC. furcillata

Olcostephanus (Olcostephanus) nicklesi Neocomites peregrinus

N. peregrinus Karakaschiceras pronecostatum

Upper

Saynoceras verrucosum S. verrucosum

Neocomites platycostatus Karakaschiceras biassalense

Saynoceras fuhri Busnardoites campylotoxus B. campylotoxus

VA

LA

NG

INIA

N

Lower

Tirnovella pertransiens Thurmanniceras otopeta

Tirnovella alpillensis Berriasella picteti

Upper Subthurmannia boissieri

Malbosiceras paramimounumDalmasiceras dalmasiBerriasella privasensis Middle Subthurmannia occitanica

Subthurmannia subalpina BE

RR

IASI

AN

Lower Berriasella jacobi

S. Reboulet and J. Klein (reporters) et al. / Cretaceous Research 30 (2009) 496–502498

2.2. Barremian

8. Taveraidiscus hugii auctorum Zone. The term ‘‘auctorum’’should be kept because this index-species is not revised and thereis not consensus on its concept. The zone is subdivided into a T.hugii auctorum Subzone below and a Psilotissotia colombianaSubzone above. P. colombiana was considered as index-species ofzone (Vermeulen, 1996), and then of subzone (Vermeulen, 2003,2005). P. colombiana, revised by Vermeulen (1997b), who defineda neotype, has been found in France and Columbia (Vermeulen,2003; Vermeulen and Klein, 2006).

9. Kotetishvilia nicklesi Zone. No change.

10. Nicklesia pulchella Zone. No change.11. Kotetishvilia compressissima Zone. It is divided into the Hol-

codiscus fallax (using classification of Vermeulen, 2007), Nicklesiadidayana (using classification of Vermeulen, 2003; Vermeulen andKlein, 2006), Heinzia communis and Subtorcapella defayae Horizons.The H. fallax Horizon was introduced by Company et al. (1995) inthe zonal scheme of the Betic Cordillera (Southeast Spain) and itsrank was elevated to subzone by Vermeulen (2007). This index-species was found in Southeast France (Vermeulen, 2003, 2005, andreferences) and in the Western High Atlas, Morocco (Companyet al., 2008). A N. didayana Horizon was recognized in the WesternHigh Atlas (Company et al., 2008) where it characterizes a level in

Table 2Ammonite Standard zonation of the Barremian-Albian Stages. The numbers in this Table refer to the numbers in the text.

STAGES ZONES SUBZONES HORIZONS S. (S.) dispar

Stoliczkaia (Stoliczkaia) dispar Stoliczkaia (Faraudiella) blancheti Upper

Mortoniceras inflatum Diploceras cristatum

Euhoplites lautus Euhoplites loricatus

Hoplites spathi Middle

Hoplites dentatus Lyelliceras lyelli

Douvilleiceras mammillatum

AL

BIA

N

LowerLeymeriella tardefurcata Hypacanthoplites jacobi

Upper Acanthohoplites nolani Diadochoceras nodosocostatum

Parahoplites melchioris Epicheloniceras buxtorfiEpicheloniceras gracile

Middle Epicheloniceras martiniEpicheloniceras debile

Dufrenoyia furcata Deshayesites grandis

Deshayesites deshayesi

Deshayesites weissi (16)

APT

IAN

Lower

Deshayesites oglanlensis

Pseudocrioceras waagenoides

Martelites sarasini (15)Leptoceratoides puzosianum

Heteroceras emerici

Imerites giraudi (15)

I. giraudi (15)

I. giraudi Hemihoplites feraudianus (14)

Gerhardtia provincialis Gerhardtia sartousiana (14)G. sartousiana

Barrancyloceras barremense

Upper

Toxancyloceras vandenheckii (13)T. vandenheckii (13)

Heinzia caicedi (12)Moutoniceras moutonianum (12)

Coronites darsi (12)Subtorcapella defayae (11)

Heinzia communis (11)Nicklesia didayana (11)

Kotetishvilia compressissima

Holcodiscus fallax (11)Nicklesia pulchella (10)Kotetishvilia nicklesi (9)

Psilotissotia colombiana (8)

BA

RR

EM

IAN

Lower

Taveraidiscus hugii auctorumT. hugii auctorum (8)

S. Reboulet and J. Klein (reporters) et al. / Cretaceous Research 30 (2009) 496–502 499

the middle part of the zone. This index-species is well known inSoutheast France (Vermeulen, 2003, and references). According toVermeulen (1998, 2003), a horizon of H. communis was recognizedin the upper part of the K. compressissima Zone (Hoedemaeker and

Rawson, 2000). However, this horizon was not indicated in the Lyonand Neuchatel reports (Hoedemaeker et al., 2003; Reboulet et al.,2006); probably an omission when the tables were made. H. com-munis is present in the Western High Atlas (Company et al., 2008). A

S. Reboulet and J. Klein (reporters) et al. / Cretaceous Research 30 (2009) 496–502500

S. defayae Horizon was introduced by Vermeulen (2003) andelevated to the rank of subzone (Vermeulen, 2005, 2007).

12. Moutoniceras moutonianum Zone. It replaces the formerCoronites darsi Zone sensu Vermeulen (1997a, 1998), who definedthis zone as the interval between the first appearance of C. darsi andthat of Heinzia sayni (the H. sayni Zone of Vermeulen (1997a, 1998)is equivalent to the Toxancyloceras vandenheckii Zone, see also Fig. 4in Rawson et al., 1999). The C. darsi Zone was incorporated into theStandard zonation (Hoedemaeker et al., 2003) to replace the M.moutonianum Zone which was defined by the first appearance of itsindex-species and that of T. vandenheckii (Company et al., 1995).Taking into account the data of Vermeulen (2005) and Companyet al. (2008), the first occurrence of M. moutonianum virtuallycoincides with that of C. darsi. Jean Vermeulen agrees if we considerthat M. gr. moutonianum is true M. moutonianum (see their ranges,Vermeulen, 2005, Fig. 81). In this case, the M. moutonianum (sensuCompany et al., 1995) and C. darsi (sensu Vermeulen, 1997a, 1998)Zones span the same stratigraphic interval. Consequently, andfollowing to the rules of priority (International Stratigraphic Guide,Salvador, 1994), M. moutonianum must be chosen as index-specieswhich is relatively well known (see synonymy in Klein et al., 2007).Moreover, according to some authors, the choice of M. moutonianumpresents more advantages than that of C. darsi for characterizingthis interval: a) M. moutonianum is generally more abundant(Company et al., 2008; Busnardo’s comments); b) its stratigraphicrange is longer (Company et al., 1995, 2008; Vermeulen, 2005), andc) its palaeobiogeographic distribution is wider (Klein et al., 2007;Hoedemaeker’s comments).

The M. moutonianum Zone is subdivided into a C. darsi Horizonbelow and a Heinzia caicedi Horizon above. Following Vermeulen(1997a, 1998), a horizon of H. caicedi was recognized in the Stan-dard zonation (Hoedemaeker and Rawson, 2000). However, thishorizon was not indicated in the following reports of Lyon andNeuchatel (Hoedemaeker et al., 2003; Reboulet et al., 2006),probably by an omission when the tables were made. H. caicedi hasbeen found in the Western High Atlas, Morocco (Company et al.,2008) where this species is quite common in the middle part of thezone, characterizing a level identified in most of the sectionsstudied. H. caicedi is nearly restricted to shallow palaeoenviron-ments (Vermeulen, 1998, 2003; Company et al., 2008).

13. Toxancyloceras vandenheckii Zone. According to Rawsonet al. (1996), the base of the Upper Barremian should be definedby the base of the T. vandenheckii Zone. These authors underlinedthat the index-species has a wide palaeobiogeographic distribu-tion in the Western Tethys, recorded from Spain to Georgia (seealso Klein et al., 2007). The recommendation of the 1995 Brusselsmeeting was adopted by Ogg et al. (2004). Bert et al. (2008; seealso Reboulet et al., 2007) are in favour to suppress the Holco-discus uhligi Zone (Vermeulen, 2003, 2005; Reboulet et al., 2006),and to place the Upper Barremian boundary at the base of the T.vandenheckii Zone, an index-species with a large distribution andrepresented in deep as well as in shallow palaeoenvironments.The T. vandenheckii Zone is characterized by a diversification andan important ammonoid fauna turnover more essentially withinthe Ancyloceratoidea and Hemihoplitidae with the appearance ofthe genera Toxancyloceras, ‘‘Barrancyloceras’’, Gassendiceras,Camereiceras (Bert et al., 2008), Ewaldiceras and Leroyceras(Vermeulen, 2003, 2006). Bert et al. (2008) underline that H.uhligi is hardly known (see synonymy in Klein, 2005) and itwould be useful to make a revision. The range and palae-obiogeographic distribution of this index-species should be alsobetter defined. Company et al. (2008) consider that the base of T.vandenheckii Zone marks the base of the Upper Barremian.Consequently, most of members of the Kilian Group (see alsoBusnardo in Reboulet et al., 2007; Hoedemaeker’s comment)

agree to use the solution of the Brussels meeting (Rawson et al.,1996). For Jean Vermeulen, the base of the Upper Barremianshould be placed at the base of the H. uhligi Zone as it corre-sponds to a lithological/sequential change and to the historicalboundary (Vermeulen, 2003, 2005).

The proposal of the 2007 Digne meeting (Reboulet et al., 2007)to subdivide the T. vandenhecki Zone in H. uhligi, Heinzia sayni andBarrancyloceras barremense Subzones (see also Vermeulen andLazarin, 2007; Bert et al., 2008) is abandoned. It is subdivided intoa T. vandenhecki Subzone below and B. barremense Subzone above.The ‘‘Ancylezeiceras’’ breistrofferi and the Camereiceras marchandiHorizons proposed by Bert et al. (2008) are not included in theStandard zonation.

14. Gerhardtia sartousiana Zone. According to Bert et al. (2008;see also Reboulet et al., 2007), the former zone of Hemihoplitesferaudianus is lowered in rank of subzone and occupies the upperpart of the G. sartousiana Zone. The new boundary betweenG. sartousiana and Imerites giraudi Zones is now characterized by animportant faunal turnover (disappearance of Pulchellidae andHemihoplitinae, and proliferation of Heteroceratidae mainly rep-resented by Heteroceras), and it corresponds to a transgression(sequential change; Arnaud, 2005).

The proposal of Bert et al. (2008) to define a Camereiceraslimentinus Horizon/Subzone and Gerhardtia provincialis, Hemi-hoplites feraudianus, Hemihoplites casanovai Pseudoshasticriocerasmagnini, Pseudoshasticrioceras autrani Horizons is not followed.

15. Imerites giraudi Zone. According to Bert et al. (2008; seealso Reboulet et al., 2007), the former zone of Martellites sarasiniis lowered in rank of subzone and occupies the middle part ofthe I. giraudi Zone. For these authors, the boundary between I.giraudi Zone (sensu Reboulet et al., 2006) and the former zone ofM. sarasini does not correspond to a faunal turnover. Specimensof Martelites (Martelites sp. and Martelites aff. gamkrelidzei) havebeen found in the I. giraudi Zone (sensu Reboulet et al., 2006) ofseveral sections of the stratotype area (Barreme-Angles-Cas-tellane area, Alpes-de-Haute-provence, southeastern France). Thegenus Heteroceras is still represented by two species (Heterocerasbaylei and Heteroceras fuhrae) in the former M. sarasini Zone.Previously, Kakabadze (1983, 1989) had already noted the simi-larity of the faunal assemblages between the I. giraudi Zone andColchidites securiformis Zone (nearly equivalent to the former M.sarasini Zone) of Georgia. Thus, he had proposed to considerthem as two subzones of the same zone. Jean Vermeulendisagrees with this zonal scheme and considers M. sarasini asindex-species of a zone (see also in Reboulet et al., 2007; Ver-meulen and Lazarin, 2007).

Contrary to the suggestion of Bert et al. (2008), Pseudocrio-ceras waagenoides should be considered as an index-species ofa subzone (see also Reboulet et al., 2007; P. Ropolo’s comments).Even if its distribution is controlled by ecological factors (prox-imality versus distality; great and low abundances in shallowand deep waters palaeoenvironments, respectively; Rebouletet al., 2006), P. waagenoides has a wide distribution allowingcorrelations.

2.3. Aptian

16. Deshayesites weissi Zone. Considering that D. weissi isa nomen dubium (Reboulet et al., 2006), Naser Raisossadat suggestseither to find a type from specimen of the Schloenbach collection inBerlin (Germany) or to replace Deshayesites planus instead of D.weissi in the Standard zonation. Both species are found together.Members are in favour of the first solution and Jaap Klein will workon the Schloenbach collection.

S. Reboulet and J. Klein (reporters) et al. / Cretaceous Research 30 (2009) 496–502 501

3. Nomination of the third vice-chair andnext meeting of the Kilian Group

Taking into account that around 33% of members of the KilianGroup are from countries of Eastern Europe, the chairman suggesteda nomination for a third vice-chair in order to (a) represent the groupduring symposiums which would take place in these countries and(b) to organize/animate ‘‘informal’’ Kilian Group meetings in thispart of the world. Ottilia Szives, present at the 2008 Vienna meeting,was willing to be vice-chairwoman. As none of other attendants ofthe meeting and colleagues from Eastern countries informed later bythe chairman raised up any objection, she was appointed.

The next meeting of the Group will be in Dijon (France) inSeptember 2010, together with the 8th International SymposiumCephalopods present and past. On this planned meeting, the KilianGroup is expected to focus on the problems of the Aptian andAlbian Stages.

We thank the anonymous reviewer for comments.

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