503

Nova Hedwigia 84 3mdash4 503mdash512 Stuttgart May 2007

DOI 1011270029-503520070084-0503 0029-5035070084-0503 $ 250copy 2007 J Cramer in der Gebruumlder Borntraeger

Verlagsbuchhandlung D-14129 Berlin middot D-70176 Stuttgart

Typification of twenty-six varieties of Orthotrichum(Orthotrichaceae Bryopsida) published by G Venturi

by

Ricardo Garilleti1 Francisco Lara2 and Vicente Mazimpaka2

1Departamento de Botaacutenica Facultad de Farmacia Universidad de Valencia AvdaVicente Andreacutes Estelleacutes sn E-46100 Burjasot (Valencia) Spain

E-mail ricardogarilletiuves2Departamento de Biologiacutea (Botaacutenica) Facultad de Ciencias

Universidad Autoacutenoma de Madrid CDarwin 2 E-28049 Madrid SpainE-mail franciscolarauames vicentemazimpakauames

With 2 figures and 1 table

Garilleti R F Lara amp V Mazimpaka (2007) Typification of twenty-six varieties of Orthotrichum(Orthotrichaceae Bryopsida) published by GVenturi - Nova Hedwigia 84 503-512

Abstract The Italian bryologist Gustavo Venturi was a prestigious taxonomist of the 19th Centurywhose principal study object was the genus Orthotrichum Hedw The main results of such an interestwere the description of 20 species 3 subspecies and 42 varieties validly published within this genusThe original material on which his studies were based is currently kept at the Herbarium of theMuseo Tridentino di Scienze Naturali (TR) After a revision of the Venturirsquos collection 24Orthotrichum varieties described upon European materials together with an American variety and anAsiatic one are here typified

Keywords Orthotrichum nomenclature herbarium bryophytes

Introduction

Gustavo Venturi (Roveretto 1830 - Trento 1898) was one of the most prominentEuropean Orthotrichologists Throughout his scientific life he gathered a collectionof near 4600 specimens from all bryophyte groups collected by himself or obtainedthrough exchanges with the most important contemporaneous bryologists (BrotherusDe Notaris Britton Cardot Limpricht Schimper etc) His herbarium was donatedto the city of Trento and is part of the Museo Tridentino di Scienze Naturali (TR)since its foundation in 1922

504

The specimens of the Venturirsquos collection have been perfectly preserved They arekept in wood cases in which the sheets are included in folders alphabetically orderedby species Each sheet (Figs 1-2) consists in a cardboard on which a packet containingthe specimens (= samples) is fixed The cardboards of each species are correlativelynumbered in pencil generally in the upper or inferior right corner Identificationand collecting data (geographical origin collector donor etc) of each taxon areannotated on the cardboard sometimes incompletly so The sheets frequently containmaterials from several localities and collectors In this case the samples are directlyfixed within the packet or kept in other smaller individual packets Anyway localitiesand specimens are clearly identified each by a number (Fig 1) It is also frequent tofind drawings sometimes in colour that show relevant details of the specimen(Fig 2) Although most of them were made by Venturi himself some others attributableto JCardot and of excellent quality are also found

Orthotrichum is currently represented in the TR-Venturi herbarium by 1037 specimenswhich are kept in 400 sheets under 102 different names Throughout his scientificlife Venturi validly described 20 species 3 subspecies and 42 varieties within thegenus Orthotrichum which were published in 20 papers and collaborations Vitt ampNimis (1987) typified 26 of these names (18 species 1 subspecies and 7 varieties)Unfortunately the original material on which these typifications were based was lostin course of their postal return to TR after having been studied by JLewinsky(Lewinsky in lit) In view of the necessary typification to be made again it will notbe possible to use any material of the Venturirsquos herbarium since no duplicate of the

Figure 1 Sheet number 22 of Orthotrichum pallens Brid in the TR-Venturirsquos herbarium Thecardboard is numbered in the inferior right angle (22) and includes specimens from 3 differentlocalities identified with a roman numeral for each sample Specimens are fixed in the inside of thepacket

505

material selected for that typification nor any material susceptible to be selected asneotype is kept there

Vitt amp Nimis (1987) did not include in their work the European varieties for thereason given by these authors themselves ldquoneither of us is familiar with intraspecificvariation of Orthotrichum on the European continentrdquo Afterwards only one varietydescribed by Venturi has been typified (Garilleti et al 2002) and no further study onthe herbarium of this orthotrichologist has been carried out The present work intendsto partially fill in this gap through a typification of the European varieties whoseoriginal material is kept at the Venturirsquos herbarium (TR-Venturi) For this purpose allthe Orthotrichum material gathered by Venturi and available in TR has been revisedTaxa validly published by Venturi and not treated in the present paper are summarizedon table 1 where those already typified and those for which no typifiable material hasbeen found in TR are indicated Only two species two subspecies and eight varietiescould not be typified because of the complete lack of suitable specimens at TR

For the typification it has been considered that a specimen constitutes a holotype of aname when a direct reference to that collection is made in the protologue according tothe Article 9 Note 1 of the International Code of Botanical Nomenclature (Greuter et al2000) When it has been necessary to designate a lectotype due to the existence of morethan one specimen or to the citation of different materials in the original description thespecimen that best represents the Venturirsquos concept of the taxon was selected

Figure 2 Lectotype of Orthotrichum pallens Bruch ex Brid var crispatulum Venturi in the TR-Venturirsquos herbarium It is an example of the numerous sheets containing illustrations made by Venturihimself The fixed specimens belong to three different species More details on the lectotypificationin text

506

Table 1 Other Orthotrichum taxa validly published by Gustavo Venturi The 39 Orthotrichum taxavalidly published by Venturi and not treated in this paper are here listed The reference containingthe corresponding typification or the lack of original material in TR is indicated Neotypes shouldbe designated for the names typified by Vitt amp Nimis (1987) (see more information in text)

O affine Schrad ex Brid var densum Venturi Type not locatedHedwigia 12 35 1873O affine var pulvinatum Venturi Hedwigia 12 35 1873 Type not locatedO anomalum Hedw subsp defluens Venturi Rev Bryol 9 58 1882 Type not locatedO anomalum var americanum Venturi in Macoun Type not locatedCat Canad Pl 6 86 1892 hom illegO anomalum var ovalis Venturi in Heribaud Type not locatedMem Ac Sc Clermont-Ferrand ser 2 14 322 1899O baldacci Bottini amp Venturi Hedwigia 31 136 1892 Typified in Vitt amp Nimis (1987)O californicum Venturi Hedwigia 12 6 1873 Typified in Vitt amp Nimis (1987)O caucasicum Venturi in Husnot 1887 Tyipified in Vitt amp Nimis (1987)O cupulatum var minus Venturi Hedwigia 12 7 1873 hom illeg Type not locatedO cupulatum var winterii Venturi in Geheeb Type not locatedRev Bryol 4 19 1877O duthiei Venturi Rev Bryol 23 27 1896 Typified in Vitt amp Nimis (1987)O euryphyllum Venturi in Roell Bot Centralbl 44 417 1890 Tyipified in Vitt amp Nimis (1987)O firmum Venturi Nuov Giorn Bot Ital 4 15 1872 Type not locatedO lateciliatum Venturi in Broth Typified in Vitt amp Nimis (1987)Oefv Finsk Vet Soc Foerh 35 44 1893O lateciliatum Venturi var apiculatum Venturi Type not locatedPap Proc R Soc Tasmania 1894-95 112 1895O lyellii Hook amp Tayl subsp strictum Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O nudum Dicks var rudolphianum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O pallens Brid var parvum Venturi Hedwigia 12 21 1873 Typified in Vitt amp Nimis (1987)O philibertii Venturi Rev Bryol 5 45 1878 Type not locatedO praemorsum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O preperistomatum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pseudopumilum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pulchellum Brunt var leucodon Venturi in Roell Type not locatedBot Centralbl 44 419 1890O rhabdophorum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O rivulare Turn var dentatum Venturi in Geheeb Typified in Vitt amp Nimis (1987)Rev Bryol 23 60 1896O roellii Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O rupestre Schw var ovatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O rupestriforme Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O sardagnanum Venturi Rev Bryol 6 56 1879 Typified in Vitt amp Nimis (1987)O schlotthaueri Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O speciosum Nees var roellii Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O stenocarpum Venturi in Roell Bot Centralbl 44 389 1890 Typified in Vitt amp Nimis (1987)O stramineum Hornsch ex Brid var defluens Venturi Typified in Garilleti et al (2002)Hedwigia 12 19 1873O strangulatum PBeauv var mucronatum Venturi Type not locatedHedwigia 12 23 1873O urnigerum Myr var perforatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O venturii De Not subsp fuscum Venturi Rev Bryol 8 47 1881 Type not locatedO venustum Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O virens Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O vladikavkanum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)

507

Besides the European names an American taxon Orthotrichum pumilum Sw exanon var americanum Venturi in Husnot and an Asiatic one Orthotrichum alpestreHornsch ex Bruch amp Schimp var dilatatum Venturi in Broth which had not beenincluded in the Vitt amp Nimisrsquo work (1987) are here also typified

No typification of the many ldquotypicumrdquo varieties published by Venturi has been donesince they are invalid names in accordance with the articles 243 and 262 of theICBN In case of the varieties ldquocommunerdquo that Vitt amp Nimis (1987) consideredmore problematic than the ldquotypicumrdquo ones the same criterion has been followedthrough the application of the article 262 For this we based our criterion on Venturirsquoscomments that allow to understand what this author understood by ldquocommunerdquovarieties and show without any doubt that these varieties include the type of thespecies For instance in case of Orthotrichum pallens Bruch ex Brid var communeVenturi (1887) comments ldquoUn eacutechantillon original de lrsquoO pallens communiqueacutepar Bruch lui-mecircme agrave De Notaris appartient agrave cette varieacuteteacute [commune] dans tous sesdeacutetailsrdquo It is also the case of Orthotrichum rogeri Brid var commune since Venturi(1887) himself bases its description on the original materials of the type varietyldquodes eacutechantillons authentiques de lrsquoO rogeri de Bridel et de Schimperrdquo It is thesame case as Orthotrichum rupestre Schleich ex Schwaumlgr var vulgare Venturi sinceVenturi (1887) includes in this taxon the species type as well as O flaccum De Not

Typification

Orthotrichum affine Schrad ex Brid var medium Venturi Hedwigia 12 35 1873

TYPE Ex Anglia et Germania

LECTOTYPE Single sheet in own folder specimen labelled as Orthotrichum medium Wils msldquoBelow Underbarrow Mill Juni 1869 Davies ddt et legit 1871rdquo TR-Venturi here designated

Orthotrichum alpestre Hornsch ex Bruch amp Schimp var dilatatum Venturi inBroth Acta Soc Sci Fenn 24(2) 17 1898

TYPE Kashmir Kishenganga Valley near Gurois 791893 Duthie 13988

HOLOTYPE Sheet 7 in the folder of O alpestre specimen labelled as Orthotrichum alpestre varannotated ldquovar dilatatardquo on the packet ldquo13988 Kashmir Kischangonga valley 8000 p 79893 legJFDuthie comm Brotherusrdquo TR-Venturi

Orthotrichum anomalum Hedw var ibericum Venturi in Leresche amp Levier DeuxExcursions Botaniques 170 1880

TYPE Picos de Europa 1879

HOLOTYPE Sheet 26 in the folder of O anomalum separated in a subfolder of ldquovar ibericumrdquospecimen labelled as ldquoOrthotrichum anomalum v hibericum [sic] mihi origrdquo ldquoPico de EuropaProv Santander in alpinis editioribus 2000rsquo ad rupes calcareas Levier et Leresche lgt 15 Juli lsquo78rdquoTR-Venturi

OBSERVATION The protologue is apparently confusing on the collecting date While the reference yearof this taxon (in the protologue) is ldquo1879rdquo it is formerly established in the publication that thespecimens were collected during July and August 1878

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

504

The specimens of the Venturirsquos collection have been perfectly preserved They arekept in wood cases in which the sheets are included in folders alphabetically orderedby species Each sheet (Figs 1-2) consists in a cardboard on which a packet containingthe specimens (= samples) is fixed The cardboards of each species are correlativelynumbered in pencil generally in the upper or inferior right corner Identificationand collecting data (geographical origin collector donor etc) of each taxon areannotated on the cardboard sometimes incompletly so The sheets frequently containmaterials from several localities and collectors In this case the samples are directlyfixed within the packet or kept in other smaller individual packets Anyway localitiesand specimens are clearly identified each by a number (Fig 1) It is also frequent tofind drawings sometimes in colour that show relevant details of the specimen(Fig 2) Although most of them were made by Venturi himself some others attributableto JCardot and of excellent quality are also found

Orthotrichum is currently represented in the TR-Venturi herbarium by 1037 specimenswhich are kept in 400 sheets under 102 different names Throughout his scientificlife Venturi validly described 20 species 3 subspecies and 42 varieties within thegenus Orthotrichum which were published in 20 papers and collaborations Vitt ampNimis (1987) typified 26 of these names (18 species 1 subspecies and 7 varieties)Unfortunately the original material on which these typifications were based was lostin course of their postal return to TR after having been studied by JLewinsky(Lewinsky in lit) In view of the necessary typification to be made again it will notbe possible to use any material of the Venturirsquos herbarium since no duplicate of the

Figure 1 Sheet number 22 of Orthotrichum pallens Brid in the TR-Venturirsquos herbarium Thecardboard is numbered in the inferior right angle (22) and includes specimens from 3 differentlocalities identified with a roman numeral for each sample Specimens are fixed in the inside of thepacket

505

material selected for that typification nor any material susceptible to be selected asneotype is kept there

Vitt amp Nimis (1987) did not include in their work the European varieties for thereason given by these authors themselves ldquoneither of us is familiar with intraspecificvariation of Orthotrichum on the European continentrdquo Afterwards only one varietydescribed by Venturi has been typified (Garilleti et al 2002) and no further study onthe herbarium of this orthotrichologist has been carried out The present work intendsto partially fill in this gap through a typification of the European varieties whoseoriginal material is kept at the Venturirsquos herbarium (TR-Venturi) For this purpose allthe Orthotrichum material gathered by Venturi and available in TR has been revisedTaxa validly published by Venturi and not treated in the present paper are summarizedon table 1 where those already typified and those for which no typifiable material hasbeen found in TR are indicated Only two species two subspecies and eight varietiescould not be typified because of the complete lack of suitable specimens at TR

For the typification it has been considered that a specimen constitutes a holotype of aname when a direct reference to that collection is made in the protologue according tothe Article 9 Note 1 of the International Code of Botanical Nomenclature (Greuter et al2000) When it has been necessary to designate a lectotype due to the existence of morethan one specimen or to the citation of different materials in the original description thespecimen that best represents the Venturirsquos concept of the taxon was selected

Figure 2 Lectotype of Orthotrichum pallens Bruch ex Brid var crispatulum Venturi in the TR-Venturirsquos herbarium It is an example of the numerous sheets containing illustrations made by Venturihimself The fixed specimens belong to three different species More details on the lectotypificationin text

506

Table 1 Other Orthotrichum taxa validly published by Gustavo Venturi The 39 Orthotrichum taxavalidly published by Venturi and not treated in this paper are here listed The reference containingthe corresponding typification or the lack of original material in TR is indicated Neotypes shouldbe designated for the names typified by Vitt amp Nimis (1987) (see more information in text)

O affine Schrad ex Brid var densum Venturi Type not locatedHedwigia 12 35 1873O affine var pulvinatum Venturi Hedwigia 12 35 1873 Type not locatedO anomalum Hedw subsp defluens Venturi Rev Bryol 9 58 1882 Type not locatedO anomalum var americanum Venturi in Macoun Type not locatedCat Canad Pl 6 86 1892 hom illegO anomalum var ovalis Venturi in Heribaud Type not locatedMem Ac Sc Clermont-Ferrand ser 2 14 322 1899O baldacci Bottini amp Venturi Hedwigia 31 136 1892 Typified in Vitt amp Nimis (1987)O californicum Venturi Hedwigia 12 6 1873 Typified in Vitt amp Nimis (1987)O caucasicum Venturi in Husnot 1887 Tyipified in Vitt amp Nimis (1987)O cupulatum var minus Venturi Hedwigia 12 7 1873 hom illeg Type not locatedO cupulatum var winterii Venturi in Geheeb Type not locatedRev Bryol 4 19 1877O duthiei Venturi Rev Bryol 23 27 1896 Typified in Vitt amp Nimis (1987)O euryphyllum Venturi in Roell Bot Centralbl 44 417 1890 Tyipified in Vitt amp Nimis (1987)O firmum Venturi Nuov Giorn Bot Ital 4 15 1872 Type not locatedO lateciliatum Venturi in Broth Typified in Vitt amp Nimis (1987)Oefv Finsk Vet Soc Foerh 35 44 1893O lateciliatum Venturi var apiculatum Venturi Type not locatedPap Proc R Soc Tasmania 1894-95 112 1895O lyellii Hook amp Tayl subsp strictum Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O nudum Dicks var rudolphianum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O pallens Brid var parvum Venturi Hedwigia 12 21 1873 Typified in Vitt amp Nimis (1987)O philibertii Venturi Rev Bryol 5 45 1878 Type not locatedO praemorsum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O preperistomatum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pseudopumilum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pulchellum Brunt var leucodon Venturi in Roell Type not locatedBot Centralbl 44 419 1890O rhabdophorum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O rivulare Turn var dentatum Venturi in Geheeb Typified in Vitt amp Nimis (1987)Rev Bryol 23 60 1896O roellii Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O rupestre Schw var ovatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O rupestriforme Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O sardagnanum Venturi Rev Bryol 6 56 1879 Typified in Vitt amp Nimis (1987)O schlotthaueri Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O speciosum Nees var roellii Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O stenocarpum Venturi in Roell Bot Centralbl 44 389 1890 Typified in Vitt amp Nimis (1987)O stramineum Hornsch ex Brid var defluens Venturi Typified in Garilleti et al (2002)Hedwigia 12 19 1873O strangulatum PBeauv var mucronatum Venturi Type not locatedHedwigia 12 23 1873O urnigerum Myr var perforatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O venturii De Not subsp fuscum Venturi Rev Bryol 8 47 1881 Type not locatedO venustum Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O virens Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O vladikavkanum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)

507

Besides the European names an American taxon Orthotrichum pumilum Sw exanon var americanum Venturi in Husnot and an Asiatic one Orthotrichum alpestreHornsch ex Bruch amp Schimp var dilatatum Venturi in Broth which had not beenincluded in the Vitt amp Nimisrsquo work (1987) are here also typified

No typification of the many ldquotypicumrdquo varieties published by Venturi has been donesince they are invalid names in accordance with the articles 243 and 262 of theICBN In case of the varieties ldquocommunerdquo that Vitt amp Nimis (1987) consideredmore problematic than the ldquotypicumrdquo ones the same criterion has been followedthrough the application of the article 262 For this we based our criterion on Venturirsquoscomments that allow to understand what this author understood by ldquocommunerdquovarieties and show without any doubt that these varieties include the type of thespecies For instance in case of Orthotrichum pallens Bruch ex Brid var communeVenturi (1887) comments ldquoUn eacutechantillon original de lrsquoO pallens communiqueacutepar Bruch lui-mecircme agrave De Notaris appartient agrave cette varieacuteteacute [commune] dans tous sesdeacutetailsrdquo It is also the case of Orthotrichum rogeri Brid var commune since Venturi(1887) himself bases its description on the original materials of the type varietyldquodes eacutechantillons authentiques de lrsquoO rogeri de Bridel et de Schimperrdquo It is thesame case as Orthotrichum rupestre Schleich ex Schwaumlgr var vulgare Venturi sinceVenturi (1887) includes in this taxon the species type as well as O flaccum De Not

Typification

Orthotrichum affine Schrad ex Brid var medium Venturi Hedwigia 12 35 1873

TYPE Ex Anglia et Germania

LECTOTYPE Single sheet in own folder specimen labelled as Orthotrichum medium Wils msldquoBelow Underbarrow Mill Juni 1869 Davies ddt et legit 1871rdquo TR-Venturi here designated

Orthotrichum alpestre Hornsch ex Bruch amp Schimp var dilatatum Venturi inBroth Acta Soc Sci Fenn 24(2) 17 1898

TYPE Kashmir Kishenganga Valley near Gurois 791893 Duthie 13988

HOLOTYPE Sheet 7 in the folder of O alpestre specimen labelled as Orthotrichum alpestre varannotated ldquovar dilatatardquo on the packet ldquo13988 Kashmir Kischangonga valley 8000 p 79893 legJFDuthie comm Brotherusrdquo TR-Venturi

Orthotrichum anomalum Hedw var ibericum Venturi in Leresche amp Levier DeuxExcursions Botaniques 170 1880

TYPE Picos de Europa 1879

HOLOTYPE Sheet 26 in the folder of O anomalum separated in a subfolder of ldquovar ibericumrdquospecimen labelled as ldquoOrthotrichum anomalum v hibericum [sic] mihi origrdquo ldquoPico de EuropaProv Santander in alpinis editioribus 2000rsquo ad rupes calcareas Levier et Leresche lgt 15 Juli lsquo78rdquoTR-Venturi

OBSERVATION The protologue is apparently confusing on the collecting date While the reference yearof this taxon (in the protologue) is ldquo1879rdquo it is formerly established in the publication that thespecimens were collected during July and August 1878

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

505

material selected for that typification nor any material susceptible to be selected asneotype is kept there

Vitt amp Nimis (1987) did not include in their work the European varieties for thereason given by these authors themselves ldquoneither of us is familiar with intraspecificvariation of Orthotrichum on the European continentrdquo Afterwards only one varietydescribed by Venturi has been typified (Garilleti et al 2002) and no further study onthe herbarium of this orthotrichologist has been carried out The present work intendsto partially fill in this gap through a typification of the European varieties whoseoriginal material is kept at the Venturirsquos herbarium (TR-Venturi) For this purpose allthe Orthotrichum material gathered by Venturi and available in TR has been revisedTaxa validly published by Venturi and not treated in the present paper are summarizedon table 1 where those already typified and those for which no typifiable material hasbeen found in TR are indicated Only two species two subspecies and eight varietiescould not be typified because of the complete lack of suitable specimens at TR

For the typification it has been considered that a specimen constitutes a holotype of aname when a direct reference to that collection is made in the protologue according tothe Article 9 Note 1 of the International Code of Botanical Nomenclature (Greuter et al2000) When it has been necessary to designate a lectotype due to the existence of morethan one specimen or to the citation of different materials in the original description thespecimen that best represents the Venturirsquos concept of the taxon was selected

Figure 2 Lectotype of Orthotrichum pallens Bruch ex Brid var crispatulum Venturi in the TR-Venturirsquos herbarium It is an example of the numerous sheets containing illustrations made by Venturihimself The fixed specimens belong to three different species More details on the lectotypificationin text

506

Table 1 Other Orthotrichum taxa validly published by Gustavo Venturi The 39 Orthotrichum taxavalidly published by Venturi and not treated in this paper are here listed The reference containingthe corresponding typification or the lack of original material in TR is indicated Neotypes shouldbe designated for the names typified by Vitt amp Nimis (1987) (see more information in text)

O affine Schrad ex Brid var densum Venturi Type not locatedHedwigia 12 35 1873O affine var pulvinatum Venturi Hedwigia 12 35 1873 Type not locatedO anomalum Hedw subsp defluens Venturi Rev Bryol 9 58 1882 Type not locatedO anomalum var americanum Venturi in Macoun Type not locatedCat Canad Pl 6 86 1892 hom illegO anomalum var ovalis Venturi in Heribaud Type not locatedMem Ac Sc Clermont-Ferrand ser 2 14 322 1899O baldacci Bottini amp Venturi Hedwigia 31 136 1892 Typified in Vitt amp Nimis (1987)O californicum Venturi Hedwigia 12 6 1873 Typified in Vitt amp Nimis (1987)O caucasicum Venturi in Husnot 1887 Tyipified in Vitt amp Nimis (1987)O cupulatum var minus Venturi Hedwigia 12 7 1873 hom illeg Type not locatedO cupulatum var winterii Venturi in Geheeb Type not locatedRev Bryol 4 19 1877O duthiei Venturi Rev Bryol 23 27 1896 Typified in Vitt amp Nimis (1987)O euryphyllum Venturi in Roell Bot Centralbl 44 417 1890 Tyipified in Vitt amp Nimis (1987)O firmum Venturi Nuov Giorn Bot Ital 4 15 1872 Type not locatedO lateciliatum Venturi in Broth Typified in Vitt amp Nimis (1987)Oefv Finsk Vet Soc Foerh 35 44 1893O lateciliatum Venturi var apiculatum Venturi Type not locatedPap Proc R Soc Tasmania 1894-95 112 1895O lyellii Hook amp Tayl subsp strictum Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O nudum Dicks var rudolphianum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O pallens Brid var parvum Venturi Hedwigia 12 21 1873 Typified in Vitt amp Nimis (1987)O philibertii Venturi Rev Bryol 5 45 1878 Type not locatedO praemorsum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O preperistomatum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pseudopumilum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pulchellum Brunt var leucodon Venturi in Roell Type not locatedBot Centralbl 44 419 1890O rhabdophorum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O rivulare Turn var dentatum Venturi in Geheeb Typified in Vitt amp Nimis (1987)Rev Bryol 23 60 1896O roellii Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O rupestre Schw var ovatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O rupestriforme Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O sardagnanum Venturi Rev Bryol 6 56 1879 Typified in Vitt amp Nimis (1987)O schlotthaueri Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O speciosum Nees var roellii Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O stenocarpum Venturi in Roell Bot Centralbl 44 389 1890 Typified in Vitt amp Nimis (1987)O stramineum Hornsch ex Brid var defluens Venturi Typified in Garilleti et al (2002)Hedwigia 12 19 1873O strangulatum PBeauv var mucronatum Venturi Type not locatedHedwigia 12 23 1873O urnigerum Myr var perforatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O venturii De Not subsp fuscum Venturi Rev Bryol 8 47 1881 Type not locatedO venustum Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O virens Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O vladikavkanum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)

507

Besides the European names an American taxon Orthotrichum pumilum Sw exanon var americanum Venturi in Husnot and an Asiatic one Orthotrichum alpestreHornsch ex Bruch amp Schimp var dilatatum Venturi in Broth which had not beenincluded in the Vitt amp Nimisrsquo work (1987) are here also typified

No typification of the many ldquotypicumrdquo varieties published by Venturi has been donesince they are invalid names in accordance with the articles 243 and 262 of theICBN In case of the varieties ldquocommunerdquo that Vitt amp Nimis (1987) consideredmore problematic than the ldquotypicumrdquo ones the same criterion has been followedthrough the application of the article 262 For this we based our criterion on Venturirsquoscomments that allow to understand what this author understood by ldquocommunerdquovarieties and show without any doubt that these varieties include the type of thespecies For instance in case of Orthotrichum pallens Bruch ex Brid var communeVenturi (1887) comments ldquoUn eacutechantillon original de lrsquoO pallens communiqueacutepar Bruch lui-mecircme agrave De Notaris appartient agrave cette varieacuteteacute [commune] dans tous sesdeacutetailsrdquo It is also the case of Orthotrichum rogeri Brid var commune since Venturi(1887) himself bases its description on the original materials of the type varietyldquodes eacutechantillons authentiques de lrsquoO rogeri de Bridel et de Schimperrdquo It is thesame case as Orthotrichum rupestre Schleich ex Schwaumlgr var vulgare Venturi sinceVenturi (1887) includes in this taxon the species type as well as O flaccum De Not

Typification

Orthotrichum affine Schrad ex Brid var medium Venturi Hedwigia 12 35 1873

TYPE Ex Anglia et Germania

LECTOTYPE Single sheet in own folder specimen labelled as Orthotrichum medium Wils msldquoBelow Underbarrow Mill Juni 1869 Davies ddt et legit 1871rdquo TR-Venturi here designated

Orthotrichum alpestre Hornsch ex Bruch amp Schimp var dilatatum Venturi inBroth Acta Soc Sci Fenn 24(2) 17 1898

TYPE Kashmir Kishenganga Valley near Gurois 791893 Duthie 13988

HOLOTYPE Sheet 7 in the folder of O alpestre specimen labelled as Orthotrichum alpestre varannotated ldquovar dilatatardquo on the packet ldquo13988 Kashmir Kischangonga valley 8000 p 79893 legJFDuthie comm Brotherusrdquo TR-Venturi

Orthotrichum anomalum Hedw var ibericum Venturi in Leresche amp Levier DeuxExcursions Botaniques 170 1880

TYPE Picos de Europa 1879

HOLOTYPE Sheet 26 in the folder of O anomalum separated in a subfolder of ldquovar ibericumrdquospecimen labelled as ldquoOrthotrichum anomalum v hibericum [sic] mihi origrdquo ldquoPico de EuropaProv Santander in alpinis editioribus 2000rsquo ad rupes calcareas Levier et Leresche lgt 15 Juli lsquo78rdquoTR-Venturi

OBSERVATION The protologue is apparently confusing on the collecting date While the reference yearof this taxon (in the protologue) is ldquo1879rdquo it is formerly established in the publication that thespecimens were collected during July and August 1878

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

506

Table 1 Other Orthotrichum taxa validly published by Gustavo Venturi The 39 Orthotrichum taxavalidly published by Venturi and not treated in this paper are here listed The reference containingthe corresponding typification or the lack of original material in TR is indicated Neotypes shouldbe designated for the names typified by Vitt amp Nimis (1987) (see more information in text)

O affine Schrad ex Brid var densum Venturi Type not locatedHedwigia 12 35 1873O affine var pulvinatum Venturi Hedwigia 12 35 1873 Type not locatedO anomalum Hedw subsp defluens Venturi Rev Bryol 9 58 1882 Type not locatedO anomalum var americanum Venturi in Macoun Type not locatedCat Canad Pl 6 86 1892 hom illegO anomalum var ovalis Venturi in Heribaud Type not locatedMem Ac Sc Clermont-Ferrand ser 2 14 322 1899O baldacci Bottini amp Venturi Hedwigia 31 136 1892 Typified in Vitt amp Nimis (1987)O californicum Venturi Hedwigia 12 6 1873 Typified in Vitt amp Nimis (1987)O caucasicum Venturi in Husnot 1887 Tyipified in Vitt amp Nimis (1987)O cupulatum var minus Venturi Hedwigia 12 7 1873 hom illeg Type not locatedO cupulatum var winterii Venturi in Geheeb Type not locatedRev Bryol 4 19 1877O duthiei Venturi Rev Bryol 23 27 1896 Typified in Vitt amp Nimis (1987)O euryphyllum Venturi in Roell Bot Centralbl 44 417 1890 Tyipified in Vitt amp Nimis (1987)O firmum Venturi Nuov Giorn Bot Ital 4 15 1872 Type not locatedO lateciliatum Venturi in Broth Typified in Vitt amp Nimis (1987)Oefv Finsk Vet Soc Foerh 35 44 1893O lateciliatum Venturi var apiculatum Venturi Type not locatedPap Proc R Soc Tasmania 1894-95 112 1895O lyellii Hook amp Tayl subsp strictum Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O nudum Dicks var rudolphianum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O pallens Brid var parvum Venturi Hedwigia 12 21 1873 Typified in Vitt amp Nimis (1987)O philibertii Venturi Rev Bryol 5 45 1878 Type not locatedO praemorsum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O preperistomatum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pseudopumilum Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O pulchellum Brunt var leucodon Venturi in Roell Type not locatedBot Centralbl 44 419 1890O rhabdophorum Venturi in Roell Bot Centralbl 44 418 1890 Typified in Vitt amp Nimis (1987)O rivulare Turn var dentatum Venturi in Geheeb Typified in Vitt amp Nimis (1987)Rev Bryol 23 60 1896O roellii Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O rupestre Schw var ovatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O rupestriforme Venturi Rev Bryol 23 67 1896 Typified in Vitt amp Nimis (1987)O sardagnanum Venturi Rev Bryol 6 56 1879 Typified in Vitt amp Nimis (1987)O schlotthaueri Venturi in Roell Bot Centralbl 44 390 1890 Typified in Vitt amp Nimis (1987)O speciosum Nees var roellii Venturi in Roell Typified in Vitt amp Nimis (1987)Bot Centralbl 44 419 1890O stenocarpum Venturi in Roell Bot Centralbl 44 389 1890 Typified in Vitt amp Nimis (1987)O stramineum Hornsch ex Brid var defluens Venturi Typified in Garilleti et al (2002)Hedwigia 12 19 1873O strangulatum PBeauv var mucronatum Venturi Type not locatedHedwigia 12 23 1873O urnigerum Myr var perforatum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)O venturii De Not subsp fuscum Venturi Rev Bryol 8 47 1881 Type not locatedO venustum Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O virens Venturi in Broth Act Soc Sc Fenn 24 (2) 18 1899 Typified in Vitt amp Nimis (1987)O vladikavkanum Venturi in Husnot 1887 Typified in Vitt amp Nimis (1987)

507

Besides the European names an American taxon Orthotrichum pumilum Sw exanon var americanum Venturi in Husnot and an Asiatic one Orthotrichum alpestreHornsch ex Bruch amp Schimp var dilatatum Venturi in Broth which had not beenincluded in the Vitt amp Nimisrsquo work (1987) are here also typified

No typification of the many ldquotypicumrdquo varieties published by Venturi has been donesince they are invalid names in accordance with the articles 243 and 262 of theICBN In case of the varieties ldquocommunerdquo that Vitt amp Nimis (1987) consideredmore problematic than the ldquotypicumrdquo ones the same criterion has been followedthrough the application of the article 262 For this we based our criterion on Venturirsquoscomments that allow to understand what this author understood by ldquocommunerdquovarieties and show without any doubt that these varieties include the type of thespecies For instance in case of Orthotrichum pallens Bruch ex Brid var communeVenturi (1887) comments ldquoUn eacutechantillon original de lrsquoO pallens communiqueacutepar Bruch lui-mecircme agrave De Notaris appartient agrave cette varieacuteteacute [commune] dans tous sesdeacutetailsrdquo It is also the case of Orthotrichum rogeri Brid var commune since Venturi(1887) himself bases its description on the original materials of the type varietyldquodes eacutechantillons authentiques de lrsquoO rogeri de Bridel et de Schimperrdquo It is thesame case as Orthotrichum rupestre Schleich ex Schwaumlgr var vulgare Venturi sinceVenturi (1887) includes in this taxon the species type as well as O flaccum De Not

Typification

Orthotrichum affine Schrad ex Brid var medium Venturi Hedwigia 12 35 1873

TYPE Ex Anglia et Germania

LECTOTYPE Single sheet in own folder specimen labelled as Orthotrichum medium Wils msldquoBelow Underbarrow Mill Juni 1869 Davies ddt et legit 1871rdquo TR-Venturi here designated

Orthotrichum alpestre Hornsch ex Bruch amp Schimp var dilatatum Venturi inBroth Acta Soc Sci Fenn 24(2) 17 1898

TYPE Kashmir Kishenganga Valley near Gurois 791893 Duthie 13988

HOLOTYPE Sheet 7 in the folder of O alpestre specimen labelled as Orthotrichum alpestre varannotated ldquovar dilatatardquo on the packet ldquo13988 Kashmir Kischangonga valley 8000 p 79893 legJFDuthie comm Brotherusrdquo TR-Venturi

Orthotrichum anomalum Hedw var ibericum Venturi in Leresche amp Levier DeuxExcursions Botaniques 170 1880

TYPE Picos de Europa 1879

HOLOTYPE Sheet 26 in the folder of O anomalum separated in a subfolder of ldquovar ibericumrdquospecimen labelled as ldquoOrthotrichum anomalum v hibericum [sic] mihi origrdquo ldquoPico de EuropaProv Santander in alpinis editioribus 2000rsquo ad rupes calcareas Levier et Leresche lgt 15 Juli lsquo78rdquoTR-Venturi

OBSERVATION The protologue is apparently confusing on the collecting date While the reference yearof this taxon (in the protologue) is ldquo1879rdquo it is formerly established in the publication that thespecimens were collected during July and August 1878

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

507

Besides the European names an American taxon Orthotrichum pumilum Sw exanon var americanum Venturi in Husnot and an Asiatic one Orthotrichum alpestreHornsch ex Bruch amp Schimp var dilatatum Venturi in Broth which had not beenincluded in the Vitt amp Nimisrsquo work (1987) are here also typified

No typification of the many ldquotypicumrdquo varieties published by Venturi has been donesince they are invalid names in accordance with the articles 243 and 262 of theICBN In case of the varieties ldquocommunerdquo that Vitt amp Nimis (1987) consideredmore problematic than the ldquotypicumrdquo ones the same criterion has been followedthrough the application of the article 262 For this we based our criterion on Venturirsquoscomments that allow to understand what this author understood by ldquocommunerdquovarieties and show without any doubt that these varieties include the type of thespecies For instance in case of Orthotrichum pallens Bruch ex Brid var communeVenturi (1887) comments ldquoUn eacutechantillon original de lrsquoO pallens communiqueacutepar Bruch lui-mecircme agrave De Notaris appartient agrave cette varieacuteteacute [commune] dans tous sesdeacutetailsrdquo It is also the case of Orthotrichum rogeri Brid var commune since Venturi(1887) himself bases its description on the original materials of the type varietyldquodes eacutechantillons authentiques de lrsquoO rogeri de Bridel et de Schimperrdquo It is thesame case as Orthotrichum rupestre Schleich ex Schwaumlgr var vulgare Venturi sinceVenturi (1887) includes in this taxon the species type as well as O flaccum De Not

Typification

Orthotrichum affine Schrad ex Brid var medium Venturi Hedwigia 12 35 1873

TYPE Ex Anglia et Germania

LECTOTYPE Single sheet in own folder specimen labelled as Orthotrichum medium Wils msldquoBelow Underbarrow Mill Juni 1869 Davies ddt et legit 1871rdquo TR-Venturi here designated

Orthotrichum alpestre Hornsch ex Bruch amp Schimp var dilatatum Venturi inBroth Acta Soc Sci Fenn 24(2) 17 1898

TYPE Kashmir Kishenganga Valley near Gurois 791893 Duthie 13988

HOLOTYPE Sheet 7 in the folder of O alpestre specimen labelled as Orthotrichum alpestre varannotated ldquovar dilatatardquo on the packet ldquo13988 Kashmir Kischangonga valley 8000 p 79893 legJFDuthie comm Brotherusrdquo TR-Venturi

Orthotrichum anomalum Hedw var ibericum Venturi in Leresche amp Levier DeuxExcursions Botaniques 170 1880

TYPE Picos de Europa 1879

HOLOTYPE Sheet 26 in the folder of O anomalum separated in a subfolder of ldquovar ibericumrdquospecimen labelled as ldquoOrthotrichum anomalum v hibericum [sic] mihi origrdquo ldquoPico de EuropaProv Santander in alpinis editioribus 2000rsquo ad rupes calcareas Levier et Leresche lgt 15 Juli lsquo78rdquoTR-Venturi

OBSERVATION The protologue is apparently confusing on the collecting date While the reference yearof this taxon (in the protologue) is ldquo1879rdquo it is formerly established in the publication that thespecimens were collected during July and August 1878

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

508

Orthotrichum anomalum Hedw var montanum Venturi Hedwigia 12 8 1873

TYPE ClLorentz ex Heiligenbluth Carinthiae retulit

HOLOTYPE Sheet 24-1 in the folder of O anomalum separated in a subfolder of ldquovar montanumrdquospecimen labelled as Orthotrichum anomalum var γ montana ldquoLorentz prope Heiligenbluth Limprichtddtrdquo TR-Venturi

Orthotrichum cupulatum Hoffm ex Brid var calcareum Venturi Hedwigia 127 1873

TYPE Ex Helvetia lectum cl Geheeb misit

HOLOTYPE Sheet kept in the folder of Orthotrichum urnigerum Myr in a separated subfolderwithout name and labelled as Orthotrichum sardagnanum [binomen crossed out] Syn Orthotrichumurnigerum var γ calcareum mihi olim ldquoSchwaitz an Kalkelsen auf dem Stoos am VienvoldstaumlthorseeMatzler lgt alt 4000rsquo Geheeb ddt 1872rdquo TR-Venturi

Orthotrichum diaphanum Schrad ex Brid var aquaticum Davies ex VenturiHedwigia 12 39 1873

TYPE Ex Anglia cum Orth spruceo commixtum possideo

HOLOTYPE Sheet 16 in the folder of O diaphanum separated in a subfolder of ldquovar inundatumrdquospecimen labelled as Orthotrichum diaphanum Schrad var inundatum Davies ldquoIt grows on treesoccasionally under water Levens Mars 1870 GDavies ddt et legitrdquo TR-Venturi

Orthotrichum leucomitrium Bruch ex Brid var elatum Venturi Hedwigia 12 241873

TYPE In sylvis vallis Rabbi

HOLOTYPE Sheet 9 in the folder of O leucomitrium separated in a subfolder of ldquovar elatiorrdquospecimen labelled as Orthotrichum leucomitrium Schimp var b elata mihi ldquoTrentino sulle coniferea Rabbi Estate 1871rdquo TR-Venturi

Orthotrichum pallens Bruch ex Brid var crispatulum Venturi Hedwigia 12 211873

TYPE In monte Baldo et in sylvis Rabbi

LECTOTYPE Sheet 29 of O pallens in subfolder of ldquovar crispulumrdquo left-sited cushion specimenlabelled as Orthotrichum pallens var b crispula Schimp ldquoSu gli alberi a Rabbi Agosto 1870Trentinordquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 27 in the folder of O pallens specimen labelled as Orthotrichumpallens var b crispula mihi ldquoTrentino corteccia di faggia Monte Baldo Agosto rsquo71 var con foliecrisperdquo TR-Venturi

OBSERVATIONS A mixture of Orthotrichum pallens and O rogeri Brid is conserved in the sheet 29 Theonly available pure cushion has been selected as the lectotype The male plants of Orthotrichum rogeriare the responsible of the crisped aspect of the cushions which seems to be teratological The onlypeculiarity of the specimen corresponding to O pallens is the frequent presence of two sporophytes ineach perichaetium The description of this variety is thus based on mixed material of two species

Orthotrichum pumilum Sw ex anon var americanum Venturi in HusnotMuscologia gallica 180 49 f 15a 1887

TYPE Sur lrsquoeacutecorce des arbres en Ameacuterique communiqueacutee par MAustin dans les Musci Appalachiani

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

509

HOLOTYPE Sheet 12 in the folder of O pumilum separated in an unidentified subfolder specimenlabelled as O pumilum americanum ldquoAustin Musci appalachiani 165 Hab on shade-trees in thetown and villages of Central New Jerseyrdquo TR-Venturi

OBSERVATIONS Material mixed with some other belonging to Orthotrichum stellatum Brid thoughspecimens corresponding to the Venturirsquos taxon dominate

Orthotrichum pumilum Sw ex anon var molle Venturi in Husnot Muscologiagallica 180 49 f 15b 1887

TYPE Sur les troncs drsquoarbres agrave Alenccedilon (Orne)

HOLOTYPE Single sheet in the subfolder of Orthotrichum rogeri [sensu Boulay] specimen labelledas Orthotrichum rogeri ldquoHusnot Musci Galliae 264 Orthotrichum rogeri BridSch suppO pallens BruchSur les arbres du champ de foire drsquoAlenccedilon (Orne) - Juinrdquo Annonation byVenturi ldquoRecte est Orth pumilum Swrdquo TR-Venturi

OBSERVATIONS This sheet is explicitely cited by Venturi in the protologue

Orthotrichum pumilum Sw ex anon var obscurum Venturi in Husnot Muscologiagallica 180 49 f 12 16 1887

TYPE M Schimper mrsquoa communiqueacute cette varieteacute dans les environs de Vienne en Autriche

HOLOTYPE Sheet 11 in the folder of O pumilum separated in an own unidentified subfolderspecimen labelled as Orthotrichum pallens ldquoPrope Vindobonensis Juratzka legt originalePhSchimper ddt rsquo79rdquo annotated ldquoO pumilum brunescensrdquo With drawings and joint notes TR-Venturi

OBSERVATIONS Although the material has a posterior annotation that identifies it as Orthotrichumpallens it completly corresponds to the Venturirsquos concept of this variety The collecting data coincidewith those given in the protologue including the lack of identification of the material received fromSchimper

Orthotrichum rupestre Schleich ex Schwaumlgr var alpinum Venturi Hedwigia 124 1873

TYPE In rupium fissuris alpis Saent In regione montana vallis Rabbi

LECTOTYPE Sheet 2-1 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varalpina ldquoAlpe Saent Sulle rupi piugrave esposte Agosto 1863rdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 2-2 in the folder of O rupestre labelled as Orthotrichum rupestrevar alpina ldquoRabbi in fissuri rupium Campisol 1871rdquo (TR-Venturi) Sheet 3 in the folder ofO rupestre labelled as Orthotrichum rupestre var β alpina mihi ldquoTrentino nella selva di RabbiSullo schisto 1871rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var boreale Venturi Hedwigia 125 1873

TYPE In Norvegia a cl Lorentz anno 1868 lectum

HOLOTYPE Sheet 26 in the folder of O rupestre specimen labelled as Orthotrichum rupestre Schlvar communis mihi ldquoChristiansund Norveg Un itin crypt PGLorentz Aug 1868rdquo TR-Venturi

OBSERVATIONS Although the label is not clear whether this is the material used by Venturi to describethe variety boreale it is however the only material whose origin collector and date coincide withthose given in the protologue The material fits the Venturirsquos description

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

510

Orthotrichum rupestre Schleich ex Schwaumlgr var elongatum Venturi Hedwigia12 4 1873

TYPE In sylvis Rabbi ad saxa

HOLOTYPE Sheet 7 in the folder of O rupestre specimen labelled as Orthotrichum rupestre var celongata mihi ldquoTrentino Nei boschi di Somrabi Valle di Rabbi Agosto 1870rdquo TR-Venturi

Orthotrichum rupestre Schleich ex Schwaumlgr var ibericum Venturi in Leresche ampLevier Deux Excursions Botaniques 171 1880

TYPE Sierra de Guadarrama Pico drsquoArvas

LECTOTYPE Sheet 4-1 in the folder of O aetnense specimen labelled as Orthotrichum sturmii ldquoInAsturiis Pico de Arvas supra Puerto de Leitariegos in graniticis alpinisrdquo TR-Venturi here designated

OTHER ORIGINAL MATERIAL Sheet 4-2 in the folder of O aetnense specimen labelled as Orthotrichumrupestre [specific epithet crossed out] aetnense ldquoSierra de Guadarrama Catal scpt ad moles graniticusmontis Pennalara supra La Granja Levier et Leresche Lgt 22 jul 1879rdquo

OBSERVATIONS Althoug conserved by Venturi as Orthotrichum aetnense there is no doubt that itcorresponds to the original material of var ibericum of O rupestre These are the only materials inthe Venturirsquos Herbarium for which localities and collectors coincide with the statement in the protologueand fit the Venturirsquos concept of this variety Later Venturi (1887) considered that the materials fromSierra de Guadarrama were identical to var aetnense (De Not) Venturi On the other hand thespecimen here selected as lectotype was initially identified as Orthotrichum sturmii which is coincidentwith the indications given in the protologue

Orthotrichum rupestre Schleich ex Schwaumlgr var montanum Venturi Hedwigia12 5 1873

TYPE Ex Helvetia et ex compluribus locis alpium

LECTOTYPE Sheet 8 in the folder of O rupestre specimen labelled as Orthotrichum rupestre varmontana mihi ldquoTrentino In sylvis Rabbi frequensrdquo TR-Venturi here designated

Orthotrichum schubartianum Lorentz var caespitosum Venturi Revue Bryolo-gique 6 8 1879

TYPE Scale di Venezia et lrsquoexemplaire du Val Mala donneacute par Carestia

LECTOTYPE Sheet 3-1 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumumigerum [specific epithet crossed out] schubartianum var caespitosa mihi ldquoDalle vette di ValMala sulla roccia dioritica presso la capanna dallrsquoalpe Cramisegravei Carestia raccolse 12 Giugno 1877rdquoTR-Venturi here designated

OBSERVATIONS On page 7 of the protologue Venturi reports the material on which this species isbased

Orthotrichum stramineum Hornsch var intermedium Venturi in HusnotMuscologia gallica 183 pl 50 f 16 17 1887

TYPE Dans les environs de Genegraveve

HOLOTYPE Sheet 21 in the folder of O stramineum specimen labelled as Orthotrichum stramineumintermedium ldquoMont Saleve Tronc de Fagus Guinet lgt lsquo86rdquo TR-Venturi

Orthotrichum stramineum Hornsch var firmum Venturi Hedwigia 12 19 1873

TYPEEx Germania obtinui

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

511

LECTOTYPE Sheet 20-1 in the folder of O stramineum separated in a subfolder of ldquovar firmumrdquospecimen labelled as Orthotrichum stramineum Schimp var α normalis et β firma mihi ldquoBelgenam su auf Steine Ruthe var brdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var saxatile Venturi Hedwigia 12 2 1873

TYPE Ex Anglia specimen originale a clar Davies accepi

HOLOTYPE Sheet 12 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var asaxatilis ldquoBarmouth Northwales GDavies ddt 1871rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var septentrionale Venturi Hedwigia12 3 1873

TYPE Ex Anglia Germania septentrionali et Norvegia habeo

LECTOTYPE Sheet 13 in the folder of O rupestre specimen labelled as ldquoOrthotrichum sturmii varseptentrionalisrdquo ldquoRabenhorst Bryotheca europaea 1008 Selchow in der NM (Preussen) legitRRutherdquo TR-Venturi here designated

Orthotrichum sturmii Hoppe amp Hornsch var variegatum Venturi Hedwigia 12 31873

TYPE Ex Norvegia a clar Lorentz lecta habeo

HOLOTYPE Sheet 5 in the folder of O sturmii specimen labelled as Orthotrichum sturmii var bvariegatum mihi ldquoChristiansund Norvegia legit Lorenz 1868rdquo TR-Venturi

Orthotrichum sturmii Hoppe amp Hornsch var virens Venturi Hedwigia 12 3 1873

TYPE Ad saxa schistosa vallis Pinegrave prope Nogareacute necnon in montanis Bolzani

LECTOTYPE Sheet 33 in the folder of O rupestre specimen labelled as Orthotrichum sturmii var cvirens ldquoTrentino Sulle pietra campestri a Nogareacute 1872rdquo TR-Venturi here designated

Orthotrichum tenellum Bruch ex Brid var decipiens Venturi in Husnot Muscologiagallica 186 1887

TYPE Plusieurs endroits du Tyrol meacuteridional et de lrsquoAllemagne

LECTOTYPE Sheet 14 in the folder of O tenellum separated in a unidentified subfolder specimenlabelled as Orthotrichum tenellum ldquoTrentino Corteccia di olivi Avio Primavera 1875rdquo TR-Venturihere designated

OBSERVATIONS This sheet has been selected as the lectotype since besides fitting the Venturirsquosconcept of the variety the packet included another label identifying it as ldquoOrth tenellum decipiensrdquo

Orthotrichum urnigerum Myrin var laxum Venturi Revue Bryologique 8 47 1881

TYPE Alpes de Aosta

HOLOTYPE Sheet 3-2 in the folder of O schubartzianum [sic] specimen labelled as Orthotrichumurnigerum [crossed out] schubartianum var caespitosa mihi ldquoAlagna-Valsesia 25 Maggio rsquo80 InBiscina N328 Carestia raccolserdquo TR-Venturi

Orthotrichum venturii De Not var caespitosum Venturi Revue Bryologique 847 1881

TYPE Environs du Monte-Gavia (eacutetiqueteacute par MLorentz lui-mecircme comme le preacuteceacutedent [O schu-bartianum Lorentz])

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006

512

HOLOTYPE Sheet 6-1 in the folder of O schubartzianum [sic] specimen labelled on the packet asOrthotrichum venturii caespitosus ldquoCol di Gavia 1 Exemplar originalerdquo TR-Venturi

OBSERVATIONS Though collected and distributed by Lorentz himself (ldquoExemplar originalerdquo) Venturi(1881 43) justifies why this sheet does not correspond to Orthotrichum schubartianum but it hasthe basic characters of O venturii This specimen is directly referred to by Venturi (1881 45) as thematerial on which this new variety has been based

Acknowledgments

We gratefully acknowledge the collaboration and facilities given by FTisi curator of the herbariumTR to one of the authors (FL) during his stay in Trento for the study of the Venturirsquos herbarium TheUniversidad Autoacutenoma de Madrid is also acknowledged for funding that stay This work has beenalso funded by the Spanish Ministry of Education and Science grant CGL2004-03513

References

GARILLETI R LARA F amp V MAZIMPAKA (2002) New discriminatory characters forOrthotrichum rogeri Brid (Orthotrichaceae Bryopsida) - Nova Hedwigia 75 207-216

GREUTER W MCNEILL J BARRIE FR BURDET H-M DEMOULIN V FILGUEIRASTS NICOLSON DH SILVA PC SKOG JE TREHANE P TURLAND NJ amp DLHAWKSWORTH (2000) International Code of Botanical Nomenclature (St Louis Code) - KoeltzScientific Books Koumlnigstein [Regnum Veg vol 138]

VENTURI G (1881) Des Orthotricha urnigera - Revue Bryologique 8 41-47

VENTURI G (1887) Orthotrichum - In Husnot T 1884-1890 Muscologia gallica 154-196THusnot Orne amp FSavy Paris

VITT DH amp PL NIMIS (1987) Typification of some orthotrichaceous names published byGVenturi (Orthotrichaceae Bryopsida) - Taxon 36 108-112

Received 1 June 2006 accepted in revised version 5 July 2006