Guidelines for pest management research to reduce stored ...
Cotton Pest Management
Transcript of Cotton Pest Management
A Term Paper on
INTEGRATED MANAGEMENT OF INSECTPEST OF COTTON
Submitted toAsst. Prof. Sundar Tiwari Department of Entomology
Agriculture and Forestry UniversityRampur, Chitwan
Submitted bySushil NyaupaneENT-05M-2013
Agriculture and Forestry University
Introduction
Cotton is a soft, fluffy staple fiber that grows in a boll, or
protective capsule, around the seeds of cotton plants of the
genus Gossypium The fiber is almost pure cellulose. Under
natural conditions, the cotton bolls will tend to increase the
dispersion of the seeds. Cotton has high economic and
industrial value. The history of cotton production in Nepal
has started from 1972-73 in Banke district in the form of
small research and now this farming is being widespread in
eastern Terai part of Nepal.
Insect pests that feed on plant structures that directly
produce yield, such as growing tips and fruiting structures,
are generally the greatest problem in a cotton crop. These
pests include Helicoverpa and mirids. The cotton plant has an
indeterminate growing pattern (fruit are present for most of
the season), which allows some pests to develop through
several generations.
Major pests
Helicoverpa
Helicoverpa remains a major pest on conventional (non-
genetically modified) cotton. The two species -Helicoverpa
armigera and H. punctigera - are very similar in both appearance
and the damage they cause to cotton. However correct
identification of the species is important
because H. armigera has developed high levels of resistance to
insecticides.
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Species composition in the crop will be influenced by the time
of year and location. In temperate regions, the majority of
the H. armigera population overwinter from mid-March onwards and
emerge during September/October. Helicoverpa punctigera is usually
the dominant species through September, but seasonal variation
can lead to early infestations of H. armigera in some years.
Helicoverpa adults are mobile and produce large numbers of
eggs, leading to rapid build-up of populations under
favourable conditions. Survival is influenced by the number of
predators present as well as other mortality factors such as
wind, rainfall and temperature extremes.
Figure. Cotton bollworm (Helicoverpa armigera)
Damage
In cotton, all stages of plant growth may be attacked but
reproductive tissue is preferred. Seedlings can be 'tipped
out' when terminal buds are eaten. Chewing damage to squares
and small bolls may cause them to shed, and chewing damage to
maturing bolls may prevent normal development and can lead to
secondary fungal infections such as boll rot.
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Figure. Bollworm feeding on square
Figure. Bollworm larvae and boll damage
Monitoring and thresholds
Regular monitoring of the crop for the presence of larvae
and/or damage is necessary in order to make timely decisions
on control. This is especially important when targeting small
and possibly insecticide-resistant larvae.
Visual sampling is the recommended method. Check at least 30
plants or 3 separate metres of row for every 50 ha (larger
samples give more accurate estimates). Eggs are
not necessarily a good basis for thresholds as not all hatch
and very small larvae have high mortality rates. Assess
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beneficial insect numbers and note parasitised eggs and
larvae.
When making pest management decisions in cotton, insect
numbers alone may not be sufficient. Plant monitoring (fruit
load, yield and maturity) assists in decision making when pest
levels are just below threshold or when there are combinations
of pests.
Thresholds for conventional cotton depend on crop stage and
are:
• Seedling to flowering: 2 larvae/m or 1 larvae (>8 mm)/m.
• Flowering to cut-out: 2 larvae/m or 1 larvae (>8 mm)/m or
5 brown eggs/m.
• Cut-out: 15% open bolls - 3 larvae/m or 1 larvae
(>8 mm)/m or 5 brown eggs/m.
• 15%-40% open bolls: 5 larvae/m or 2 larvae (>8 mm)/m or
5 brown eggs/m.
Resistance
Helicoverpa armigera developed resistance against most
insecticides in the late 1990s. However, with the introduction
of genetically modified cotton, biopesticides and more
selective insecticides, insecticide resistance to older
chemistry has decreased in recent
years. Helicoverpa punctigera has no known resistance to any
insecticides but the use of more selective options is
encouraged to help preserve natural enemies.
In order to prevent insecticide resistance, the cotton
industry has developed the Insecticide Resistance Management
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Strategy (IRMS). This strategy is reviewed annually to delay
development of resistance of H. armigera to conventional
insecticides. The core IRMS principles include:
• rotation between chemical groups with different modes of
action
• limiting the time period during which an insecticide can
be used
• limiting the number of applications of one particular
insecticide.
Cultural control
Post-harvest cultivation (pupae busting) to reduce the
overwintering stage of Helicoverpa is one of the most important
cultural control practices available. Cultivation to a depth
of at least 10 cm will damage or disturb pupae, seal their
emergence tunnels and trap emerging moths. Cultivation also
leaves survivors open to attack by birds, mice, earwigs, and
wasp parasites.
Natural enemies
Beneficial insects can affect all Helicoverpa life stages:
Predators
• Egg:red and blue beetle, damsel bug, green lacewing,
brown lacewing and various ants.
• Larvae: glossy,brown and predatory shield bugs, bigeyed,
bug, damsel bug, assassin bug, red and blue beetle, brown
lacewing common
• Pupae: common brown earwig and wireworm larvae
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• Moth:spiders
Parasitoids
• Trichgramma spp. and Telenomus spp.
• Larva: tachinid flies and Microplitis demiltor
• Pupae: banded caterpillar parasite
Spider mites
The two-spotted mite (Tetranychus urticae) is the most numerous
mite species on cotton. Other species that may occur are bean
spider mite and strawberry spider mite. Mites live on the
underside of leaves and are difficult targets for aerial
sprays. Mites prefer more mature leaves and the highest
populations are usually found near the top of the plant on
main stem nodes 3-5 below the terminal. Heavy mite
infestations on cotton can originate from adjoining fields of
early sown corn from where they are carried by air currents.
Fig: spider mite on leaves
Damage
Mite infestations in seedling cotton rarely justify control
but may be a useful indicator of potential problems and should
be monitored closely. The first sign of damage is bronzing of
the upper leaf surface near the petiole or leaf fold. As
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numbers increase, the leaves turn red and become covered in
fine webbing, and affected leaves may dry and fall off. High
mite populations can significantly affect yield and quality of
cotton. The earlier in the season that infestations develop,
the greater the potential of yield loss and quality
downgrades.
Figure. Spider mite feeding damage can weaken and even kill
stressed plants
Monitoring and thresholds
Start sampling at seedling emergence and continue at least
weekly. At low numbers, mites are difficult to find. Mites are
often too numerous to count individually, so infestations can
be rated on a presence/absence system. In Australia, the
leaves on the third to fifth main stem node (down from the
plant terminal) are the best to sample.
The general threshold for mites for most of the growing season
is 30% of plants infected. The threshold depends more on when
mite populations begin to increase and how quickly they
increase rather than the number of mites. To estimate the
percentage of yield reduction caused by mites, a chart has
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been developed by the cotton industry for the different cotton
growing regions.
Management
• Under Australian conditions, mite populations increase
less rapidly on the okra leaf varieties than broadleaf
varieties.
• Mite infestations increase after the application of some
broad-spectrum insecticides for control of other pests.
• Control weeds within fields and along field boundaries
that serve as over-wintering sites for mites.
.
Thrips are the major mite predator in cotton. Others include
damsel bug, bigeyed bug, ladybird beetles, brown smudge bug,
apple dimpling bug, brown lacewing adults, and tangle web
spiders.
Mirids (Creonitiades spp)
An important sucking insect pest of cotton, mirids are
abundant in cotton from early to mid-season and occur
throughout the cotton growing regions of Queensland and New
South Wales Green mirids are more common than brown mirids but
both cause similar damage.
Damage
Adults and nymphs feed by piercing plant tissue and releasing
a chemical that destroys cells in the feeding zone, resulting
in the following symptoms:
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• localized leaf damage
• terminal wilting of young plants
• deformity (broom shaped)
• shedding of squares and small bolls
• damage to bolls (up to 15 days old) by causing warty
growths inside carpel which causes discolored lint
• malformed bolls, delay of maturity
• reduced lint and fibre quality.
Monitoring and thresholds
Mirids are a very mobile pest and populations can fluctuate
rapidly so sampling needs to be done every three days. The
beat sheet is the most effective means for estimating mirid
numbers. Monitor for number of mirids as well as for tip
damage and fruit retention on cotton plants. Adult cause more
damage than the first three nymphal instars while the fourth
and fifth instars do similar levels of damage as the adults.
Heavy tip damage and fruit retention levels can also have an
impact on control decisions.
Thresholds based on beat sheet sampling vary between warm and
cool areas due to the cotton plants ability to compensate for
damage.
• Warm areas. Early season: 4 mirids per metre. Mid
season: 3 mirids per metre.
• Cool areas. Early season: 2 mirids per metre. Mid
season: 1.5 mirids per metre.
Management
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• Control alternative hosts including native weeds before
crop establishment.
• Avoid broad-spectrum insecticides.
• Spray only when both insects and damage are at threshold
levels.
There is potential to use lucerne as a trap crop for mirids.
Lucerne should be established two weeks before cotton.
Research has shown that the addition of petroleum spray oils
to some chemicals improves product efficacy at lower (half)
label rates and helps preserve natural enemies. Research has
also demonstrated that the addition of salt (5g/L of water) to
dimethoate greatly improves that product's efficacy against
mirid nymphs and adults at rates as low as 33% of the
registered rate, while lessening the spray's impact on natural
enemies.
A number of beneficial species are known to feed on mirids yet
none are recognised as regulators of mirids in cotton
populations. Adults, nymphs and eggs are eaten by damsel bug,
big eyed bug, predatory shield bug and a number of spider
species.
Aphids
The cotton aphid (Aphis gossypii) is the most common aphid
pest. Green peach aphid is an occasional pest in the early
season and cowpea aphids can colonise cotton but rarely become
a problem.
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Figure. Cotton aphid in cotton leaves and cotton damage
Damage
Aphids cause early to late season damage to terminals, leaves,
buds and stems, and are known to transmit Cotton Bunchy Top
(CBT) diseases. The honeydew secreted by aphids can
contaminate lint once bolls begin to open.
Figure. Honeydew laden leaves from whitefly infestation
Monitoring and thresholds
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Aphids tend to be patchy in distribution so sample in several
locations. At each location collect 20 leaves from different
plants (one leaf per plant) and choose main stem leaves from
3-4 nodes below the terminal of the plant. Sample for winged
and wingless adults as well as nymphs. Non-winged adults
indicate that the population has settled in the crop. Aphids
prefer new growth so focus on terminals and recently expanded
leaves. Start sampling at seedling emergence until defoliation
and sample weekly. Where hotspots of aphids are evident, also
monitor cotton for CBT.
Significant yield loss can occur if aphid populations are
allowed to develop to high levels (>90 % of plants infested)
for a period of 2-3 weeks. Thresholds depend on the estimated
yield loss which, when compared to cotton price and control
costs, helps determine whether control is warranted. When open
bolls are present thresholds should be lowered to 50% of
plants infested or 10% of plants if honeydew is present to
prevent contamination of lint.
Management
• Control weeds that serve as alternative hosts for
aphids, including marshmallow, capeweed, thistles,
nightshade, bladder ketmia and thornapples.
• Ratoon and volunteer cotton should be controlled as they
are winter hosts and may also carryover CBT disease.
• Cotton aphid has developed widespread resistance to a
number of insecticides, so adhere to the cotton IRMS.
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Whitefly
Silverleaf whitefly (Bemisia tabaci B-biotype)(SLW) and Bemisia
tabaci Q-biotype are the most serious whitefly pests in cotton due
to their resistance to many insecticides and rapid
reproduction rate. Bemisia tabaci Q-biotype is a recent arrival
in Australia (2008) and currently not as widespread as SLW.
How this biotype will affect cotton compared to SLW is not yet
known. Others found in cotton are (Trialeurodes vaporiorium) and
eastern Australian native whitefly (Bemisia tabaci).
Whiteflies are found in all cotton growing areas of
Queensland. SLW is more prevalent in warm areas. Hosts include
cotton, sunflower, tomato, rockmelon, sowthistle weed. In
cotton growing areas, SLW normally start to reach pest levels
in December through to cotton harvest. While SLW are capable
of vectoring serious cotton viruses, no such viruses are
present in Australia. SLW is highly resistant to some
insecticides and has the ability to develop resistance to
newer insecticides if resistance is not carefully managed.
Figure. Adult whitefly
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Damage
SLW feed on plant sap and cause plants to wilt, drop leaves
and under severe pest pressure, plants may die. Excretion of
honeydew contaminates lint. One of the sugars in SLW honeydew
has a lower melting point than those found in aphid honeydew
and creates problems during processing. Honeydew promotes
sooty mould, which reduces potential crop yield by blocking
sunlight and reducing assimilation of nutrients for plant
growth.
Monitoring and thresholds
Sample once a week from first flower (777 degree days
(DD)) and then twice a week from peak flowering (1300 DD).
Sample from a minimum of 2 sites in your sample area (maximum
of 25 hectares). Count the number of adults per fifth node
leaf (sample one leaf per plant).
Thresholds are based on a scoring system. Score each leaf as
either un-infested (0 or 1 adult per leaf) or infested (2 or
more adults per leaf). Calculate the percentage of crop
infested and the crop age in day degrees as thresholds are
relative to crop development stages.
Management
• Control alternate weed hosts and consider planting cotton
away from other hosts such as soybeans, sunflower and
cucurbit crops.
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• Develop an area-wide management strategy with neighbours
including a tight planting window so that whitefly does
not migrate to successive plantings.
• Avoid early season broad-spectrum insecticides that kill
beneficial insects and subsequently flare whitefly
populations. Natural enemies play a vital role in the
management of SLW.
Minor pests
Thrips
Thrips including the tobacco thrip(also known as cotton
seedling or onion thrip),tomato thrip, and western flower
thrip are common early seedling pests but their damage is
usually cosmetic and rarely affects yield or earliness of
cotton.
Figure. thrip
Damage
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Nymphs and adults cause early season damage to terminals,
leaves, buds and stems. Infestations are more serious in cool
dry conditions when seedlings are unable to grow away from
damage. Severe damage results in destruction of the growing
point (tipping out) which leads to lateral branching. Seedling
death can occur due to severe infestation but this is often
associated with damping off diseases.
While considered a pest, thrips also predate on spider mite
eggs. In recent seasons western flower thrips have been
abundant on terminal growth later in the season.
Figure. damage of thrips in cotton
Monitoring and thresholds
Sample weekly from emergence until the crop has six true
leaves for number of thrips/plant, checking for both nymphs
and adults (presence of nymphs indicates that the population
is actively breeding). Check the number of thrips on 20-
30 plants for every 50 ha of crop. Look for symptoms of tip
damage which appear as extensive blackening of the small
leaves within the terminal. Adults are highly mobile and
nymphs are very small so counts may vary with the time of day.
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The damage potential also varies with crop vigour and
temperature.
For seedlings to six true leaves the threshold is 80%
reduction in leaf area and 10 thrips/plant (adults and
nymphs). Thresholds may need to be lowered in adverse
conditions such as cold weather, herbicide damage and damping
off disease.
Management
• Consider that thrips are natural enemies of spider
mites. Use chemical control only if thrips damage
exceeds the threshold.
• Cotton crops planted next to cereal crops are more at
risk of thrips infestations.
• Thrips prefer cooler temperatures and populations
decline if temperatures rise above 30ºC.
• Western flower thrips are resistant to some insecticide
groups.
Predators of thrips include pirate bug, lacewing larvae, brown
lacewing and ladybird beetle.
Sucking bugs
Green vegetable bug (Nezara viridula)
Green vegetable bug(GVB)is an emerging pest of cotton. In the
past, sprays applied for other pests have also controlled GVB,
however the adoption of genetically modified cotton and
greater implementation of IPM has resulted in reduced
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insecticide use and GVB populations are increasingly found in
cotton crops.
The crop is most susceptible to damage from flowering onwards.
Both nymphs and adults cause damage to bolls. External damage
is visible through feeding marks and internal damage shows up
as wart-like growths and lint staining. Damage to bolls older
than 18 days is uncommon with the preferred age about 12 days
and less.
Monitoring and thresholds
Crops should be inspected for GVB twice weekly in early to
mid-morning from boll set to boll maturity.Beat sheet
sampling is the most efficient monitoring method
as infestations tend to be patchy. Monitor as many sections of
the field as practical. GVB nymphs are more difficult to
sample accurately as their distribution is extremely clumped,
particularly during the early nymphal stages. Regular damage
assessment from boll set to boll maturity is recommended -
randomly select 14 day old bolls and look for the presence of
warts or stained lint. Damage symptoms cannot be distinguished
from those caused by mirids.
Thresholds depend on the sampling methods used:
• Visual sampling: 0.5 adults/m from flowering to harvest.
• Beat sheet sampling: 1 adult/m from flowering to harvest.
• Boll damage (14 day old bolls): 20% damage.
Nymphs of the GVB are less damaging than adults and their
numbers must be converted to adult covalents.
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Natural enemies
GVB eggs are frequently parasitised by a tiny introduced wasp,
the green vegetable bug egg parasite(Trissolcus basalis).
Parasitised eggs are easily recognised as they turn black. GVB
nymphs are attacked by ants, spiders and predatory bugs. Final
(fifth) instar and adult GVB are parasitised by the recently
introduced tachinid fly (Trichopoda giacomellii).
Pale cotton stainer bug (Dysdercus sidae)
Pale cotton stainer bugs generally arrive in cotton around the
time of first open boll. Adults mate soon after
arrival resultng in an expanding population of developing
nymphs. Cotton stainers are recognised as occasional pests of
cotton, but rarely cause economic damage and are often
controlled by broad-spectrum insecticides applied for other
pests. They cannot survive in temperatures above 40°C and need
free water for survival. In mild seasons they may need to be
managed especially in genetically modified cotton crops where
fewer insecticides are used.
Cotton stainers feed on developing and mature cotton seed,
resulting in loss of seed weight, oil content and seed
viability. Feeding on bolls up to two weeks old can kill
developing seed and cause boll shedding. Where bolls are not
shed, lint yield may be reduced and 'tightlock' around the
damaged seed prevents lint from fluffing out as bolls open.
Cotton stainers can continue feeding on seed throughout boll
development and also feed on the mature seed of open bolls,
creating bald patches (less lint). Yellow staining on lint may
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be evidence of watery faeces as bugs feed in the open bolls.
Pale cotton stainer bug damage is similar to that of green
vegetable bugs and includes a black spot on the outside of the
boll, warty growths inside the boll and brown discoloured lint
Monitoring
Beat sheet sampling is a suitable method to monitor bugs but
visual searching in the lower canopy compliments monitoring
methods. Sample at multiple sites as distribution is often
patchy. Stainer bugs tend to hide in the heat of the day and
are not easily observed at this time.
First instars are usually found on the soil or in the lower
canopy and tend to remain in clusters. Older nymphs can be
found in the lower to mid canopy and can be seen feeding in
open bolls. Once stainer presence is confirmed, monitor bolls
for signs of damage by cutting open bolls of varying ages to
confirm damage.
Natural enemies
The role of natural enemies of pale cotton stainer bugs has
not been studied in Australia. Natural enemies such as
tachinid flies (parasitic flies) and predatory bugs such as
assassin bugs have been reported in Africa.
Cotton harlequin bug (Tectocoris diophthalmus)
The cotton harlequin bug(also known as hibiscus harlequin bug)
feeds on many species belonging to the hibiscus plant family
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(Malvaceae), including ornamental hibiscus species and cotton.
It is sometimes a minor pest of cotton, feeding mostly on
young shoots, piercing the stems and sucking the sugar-rich
juices intended for shoot growth.
It has been known to cause the introduction of a fungus, which
rots the cotton boll. Nymphs feed on seeds in very small bolls
or in opening bolls. Generally damage occurs only in autumn.
Wireworms
Wireworms are larval stages of beetles that live in the soil
and generally feed on organic matter.True wireworms(Agrypnus
variabilis) prefer wet soil whereas the eastern false wireworm
(Pterohelaeus darlingensis) prefers drier conditions protected by
stubble and weeds.
Wireworms generally cause problems in spring especially in
newly developed fields, fallowed fields (with heavy trash
cover) and in fields following summer crops such as
soybeans. They bore into germinating seeds and chew through
young seedlings just below the soil level. Damage can occur
for up to four weeks after planting and heavy infestation
results in patchy stands that may need replanting.
Wireworms are difficult to sample as they move into the soil
when dry and only emerge after rain or irrigation, and
seedling damage is often the first indication of their
presence. Control measures must be applied before ors at
sowing. The most effective control is achieved by modern seed
dressings. In furrow bands, application of granular or liquid
insecticides are also effective.
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Red shouldered leaf beetle (Monolepta australis)
Red shouldered leaf beetle is a very minor pest of cotton,
damage is not common and usually confined to the edges of
fields. Larvae feed on plant roots while the adult beetle
chews terminal buds, leaves, squares and surface of bolls.
Leafhoppers (jassids)
Both the vegetable leafhopper (Austroasca viridigrisea) and cotton
leafhopper (Amrasca terraereginae) can affect cotton. Adult and
nymphs suck sap and inject toxins. Leafhoppers feed on the
upper surface of leaves. The damage appears as a pale stippled
effect. They occasionally damage seedlings and new growth.
Leafhoppers can transmit diseases such as viruses and
mycoplasma.
The current threshold for control through the early season is
50 leafhoppers per metre of row.
Minor caterpillar pests
A number of caterpillar pests can attack cotton plants. Most
of these pests are sporadic pests and rarely need control
unless large populations are present.Many caterpillar pests
use weeds as their main host plants before moving into cotton.
Control of these weeds will assist in minimising pest
infestation.
Rough bollworm (Earias huegeliana)
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Presence of rough bollworm in cotton is often associated with
the presence of the weed bladder ketmia, Hibiscus trionum. Direct
boll damage occurs when larvae tunnel into bolls. Larvae can
also be found in squares. Primary growing points can be
damaged by larvae tunnelling into the main stem. This can
occur at any growing stage including in seedlings.
The northern rough bollworm (Earias vittella) is of minor
importance in northern Australia, widely distributed
throughout Asia and is an important pest of cotton in India.
Cotton tipworm (Crocidosema plebejana)
Outbreaks of cotton tipworm are associated with winter and
spring rainfall and cool temperatures which favour growth of
Marshmallow weed Malva parviflora, the main host of tipworm. When
marshmallow dies off in early summer, tipworm move into
cotton.
Larvae tunnel into terminals destroying the single stem habit.
Subsequent multiple branching can lead to excess vegetative
growth at the expense of reproductive growth and crop
development can be delayed. At high numbers, squares and bolls
can be damaged.
Cotton looper (Anomis flava)
Loopers are leaf feeders. The cotton looper prefers older
leaves and defoliation of cotton progresses upwards on the
plant. Up to 80% defoliation has been recorded from large
infestations of cotton loopers.
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Pink spotted bollworm (Pectinophora scutigera)
The pink spotted bollworm is present in Central Queensland.
Larvae tunnel into large squares, flowers and bolls and often
complete their lifecycle within the one structure. Mature
larvae can also pupate on surface thrash and effective burial
of cotton crop residues reduces subsequent infestations.
The pink bollworm (Pectinophora gossypiella) is a major pest
overseas but currently restricted to north western Australia.
It could cause serious damage if it spread to eastern
Australian cotton regions.
Light brown apple moth (Epiphyas postvittana)
The host range of light brown apple moth includes many
broadleaf weeds as well as lucerne. It can be found infesting
cotton seedlings in cool seasons. Young larvae feed by tying
terminal leaves together with webbing.
Cotton leaf perforator (Bucculatrix gossypii)
Young larvae of the cotton leaf predator (first to third
instar) feed between the upper and lower leaf surface (mine)
while the fourth instar feeds directly on the leaf. The fifth
instar causes the most obvious damage by feeding on the lower
surface, leaving windows and holes in the leaf.
Cluster caterpillar (Spodoptera litura)
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Cluster caterpillar was a serious pest in Northern Australia
cotton growing areas but it is a minor pest in Queensland and
New South Wales. Young larvae skeletonise leaves at night;
larger caterpillars consume whole leaves. Caterpillars can
also destroy squares and flowers. Large infestations can lead
to larger areas being defoliated.
Lesser armyworm (Spodoptera exigua)
While often present in low numbers in young cotton, lesser
armyworm generally prefers weed hosts. Large numbers can cause
heavy losses in seedling crops by defoliating plants and re-
sowing may be necessary. Mature larvae can be mistaken
for Helicoverpa.
Cutworms (Agrotis spp.)
Cutworms attack cotton seedlings at or above ground soil
level. They feed in the late afternoon and at night, and can
destroy seedling by chewing through stems or eating entire
leaves.
Figure. Cutworm feeding damage
Summary & Conclusion
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Cotton, the “white gold” is one of the important commercial
crops playing a key role in the economic and industrial role.
Cotton is a soft, fluffy staple fiber that grows in a boll, or
protective capsule, around the seeds of cotton plants of the
genus Gossypium The fiber is almost pure cellulose. There are
many pests that are reducing the production and quality of the
cotton and categorized as major pests and minor pests. The
major pests of cotton include helicoverpa, spider mites,
cotton aphid and whitefly and minor pests include thrips,
green vegetable bug, cotton stainer bug, wireworm , cotton
harlequin bug, leafhopper, red shouldered leaf beetle, cutworm
etc. In this article, the description about insects, damage
symptoms and management of insect pest was included.
References
Fenemore.P.G and Prakash A.2006 2nd edition, applied
entomology., New age international P limited, publishers.
New delhi, India.
GC, Y.D.2010. Scarabs of Nepal and their microbial control.
Scarabs of Nepalese Agriculture and Microbial Control.
Lambert Acad. Publishing,Germany.
Srivastava K. P.1993., A textbook of Applied Entomology.,
Kalyani publishers., New Delhi.
Thapa, R.B., and S. Tiwari, Practical Manual of Insect Pest
Management.
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