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208 Marcin S. Przybyła, Mating systems in prehistoric populationsDOI 10.1515/pz-2013-0007 Praehistorische Zeitschrift 2013; 88(1–2): 208–225
I. Abhandlungen
Marcin S. Przybyła
Mating systems in prehistoric populations.An evolutionary approach and archaeologicalevidence
Abstract: Der aktuelle Forschungsstand deutet darauf
hin, dass die Vielfältigkeit der Heiratsformen, die sich in
vormodernen Gesellschaften beobachten lassen, das Er-
gebnis evolutionären Drucks sind. Dieser Ansicht zufolge
können Polygamie und Monogamie als konkurrierende
Strategien angesehen werden, die sich unter bestimmten
Umständen ausbreiten. Der folgende Beitrag will dies auf
der Grundlage archäologischer Daten überprüfen. Dazu
werden aus zwölf Gräberfeldern der Kupfer- und Bronze-
zeit in Mitteleuropa detaillierte Informationen zum Ge-
schlechterverhältnis und zu Unterschieden im Reichtum
der Grabausstattung ausgewertet. Diese Ergebnisse wer-
den dann mit den Vermögensunterschieden und der Sub-
sistenzgrundlage der ausgewählten Populationen ver-
glichen. Es scheint, dass diesen beiden Faktoren für die
Erklärung der Vielfalt menschlicher Heiratssysteme eine
Schlüsselrolle zukommt.
Die potentiellen Ergebnisse dieser Studie steuern
neue Argumente für die Diskussion um Heiratsformen in
frühen europäischen Gesellschaften bei. Andererseits
kann ein Vergleich gewisser Elemente archäologischer
Quellen mit Konzepten aus der Humanökologie zu einem
besseren Verständnis der Funktionsweise prähistorischer
Gesellschaften beitragen.
Keywords: Europäische Vorgeschichte; Kupferzeit; Bron-
zezeit; Heiratssysteme; Heiratsstrategien; darwinistische
Archäologie; Gräberanalyse
Abstract: L’état de la recherche actuel montre que la mul-
tiplicité des formes de mariage observées dans les sociétés
pré-modernes résulte de la pression exercée par l’évolu-
tion. De ce point de vue, la polygamie et la monogamie
peuvent être considérées comme des stratégies concur-
rentielles qui peuvent s’étendre sous certaines conditions.
Cette contribution a pour but de le vérifier sur la base de
données archéologiques. On a analysé à cet effet des infor-
mations détaillées concernant la proportion des sexes et
les écarts de richesse que présentent les mobiliers funérai-
res de douze nécropoles du Chalcolithique et de l’âge du
Bronze en Europe centrale. Les résultats obtenus sont
alors comparés aux écarts affichés par les richesses maté-
rielles et aux moyens de subsistance des populations sé-
lectionnées. Ces deux facteurs semblent jouer un rôle clé
dans l’explication de cette multitude de systèmes matri-
moniaux.
Les résultats potentiels de cette étude fournissent de
nouveaux arguments aux discussions sur les formes de
mariage dans les sociétés primitives de l’Europe. D’autre
part, la comparaison de certains éléments de sources ar-
chéologiques avec des concepts appartenant à l’écologie
humaine peut contribuer à mieux saisir la manière dont ils
reflètent le mode de fonctionnement des sociétés préhisto-
riques.
Keywords: préhistoire européenne; Chalcolithique; âge
du Bronze; systèmes matrimoniaux; stratégies matrimo-
niales; archéologie darwinienne; analyse des tombes
Abstract: The current state of research suggests that the
wide diversity in marriage forms observed in pre-modern
societies can be explained as a result of evolutionary
pressure. According to this view, both polygyny and mon-
ogamy may be considered as competing strategies which
will spread under certain circumstances. The objective of
this paper is to verify this supposition based on archae-
ological evidence. For this purpose I will examine detailed
information concerning sex ratio and differences in the
wealth of grave goods in twelve cemeteries of the Central
European Copper and Bronze Age. Then I will compare the
results with data related to the degree of wealth inequality
and the form of subsistence in the examined populations.
It seems that both factors play a key role in the origin of
variation in human mating systems.
The preliminary results of this investigation can pro-
vide new arguments in the discussion on diversity in mar-
riage forms among ancient Europeans. On the other hand,
the comparison of certain elements of archaeological evi-
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Marcin S. Przybyła, Mating systems in prehistoric populations 209
dence and concepts of human behavioural ecology can
contribute to a better understanding of the actual func-
tioning of past populations.
Keywords: European prehistory; Copper Age; Bronze Age;
mating systems; marriage strategies; Darwinian archaeol-
ogy; burial analysis
Dr. Marcin S. Przybyła: Instytut Archeologii, Uniwersytet Jagiel-loñski, ul. Gołèbia 11, 31–007 Kraków, Polska. E-Mail: [email protected]
The evolutionary study of matingsystemsMating systems in contemporary human communities are
characterised by a diversity unparalleled among other
species. It is therefore no surprise that the reasons of this
diversity have long been a subject of anthropological in-
vestigation. Studies of contemporary traditional societies,
described in the Standard Cross-Cultural Sample (SCCS)
database, have indicated that polygyny occurs among as
many as 83 % of the groups described there, although only
in 28 % of populations is the share of polygynous mar-
riages higher than 40 %. Monogamy, with a restrictive ap-
proach towards cases of polygyny, has been recorded in
17 % of these communities, and polyandry happens mar-
ginally1.
Comparative anthropological studies provide several
interesting facts which may suggest possible sources of
polygyny. One of the most frequently mentioned is a dis-
proportion in the number of men and women2. It is caused
by high male mortality rates (due to natural causes or in
accidents and warfare) and by emigration of the male part
of a population resulting from overpopulation and in-
creased rivalry for resources3. Other authors put more
stress on the role of the natural environment and environ-
mentally-determined cultural factors. Rich areas with no
geographical barriers are able to sustain large polyga-
mous families and favour mobility and migration of indi-
viduals (especially women) between groups. That is why
polygyny frequently occurs in pastoral communities, where
1 E.g. Low 1988; Ember/Ember/Low 2007, for the recent discussion
about diversity in marriage forms among historic and modern Indo-
European-speaking peoples, see Fortunato 2011.
2 Recently: Ember/Ember/Low 2007, 430–437.
3 E.g. Sherratt 1972, 501.
mobility usually goes together with aggressiveness and,
as a result, favours gaining wives via military expeditions.
Monogamy is more common among farmers and fisher-
men, permanently tied to the small territory they occupy4.
A new approach to the differentiation of mating sys-
tems has been proposed by evolutionary studies which
perceive monogamy and polygyny as the strategies
adopted by individuals to enhance their reproductive suc-
cess. For several years it has been accepted that for hu-
mans, like for other mammals5, a typical mating system is
polygyny6. Monogamy was seen as one of the cultural be-
haviours developed during the early stages of evolution in
order to reduce intra-group tensions7. In this context, it is
worthwhile mentioning the examples, known from his-
torical accounts, of polygyny among the highest social
classes of societies that otherwise practised monogamy
for cultural (e.g. religious) reasons8 (Fig. 1). However, new
empirical data have revealed no statistically significant
difference in reproductive success of men from monog-
amous and polygynous communities9. In other words,
men, unlike males of numerous other species, show no
straightforward correlation between the number of female
partners and their reproductive success10. Therefore, mon-
ogamy and polygyny may be regarded as competing strat-
egies, which will be evolutionary stable under certain spe-
cific circumstances.
These circumstances are determined on the one hand
by different interests of males and females, and on the
other hand by differences in the scale and form of trans-
mitting resources from parents to children, resulting from
the economic strategies adopted11. From a female’s point
of view, polygyny may become a preferred behaviour
when her acceptance of a partner who already has other
wives brings higher reproductive benefits than staying in a
monogamous relation. Such situation – called the “poly-
gyny threshold”12 – may occur if two factors are present: a
significant differentiation in the distribution of resources,
and their considerable impact on the reproductive success
of the offspring13.
4 White 1988; White/Burton 1988.
5 Trivers 1972.
6 E.g. Shennan 2002, 184–185; Low 2005, 68; Ember/Ember/Low
2007, 428–429.
7 Bowles/Choi/Hopfensitz 2003, 135; Alexander 2009, 71–72; 142.
8 E.g. Scheidel 2009.
9 Fortunatto/Archetti 2010, 150.
10 Brown/Laland/Borgerhoff Mulder 2009, 301–303.
11 Bowles/Smith/Borgerhoff Mulder 2010; Smith et al. 2010a; Shen-
nan 2011.
12 Orians 1969.
13 Borgerhoff Mulder 1990, 255–256.
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210 Marcin S. Przybyła, Mating systems in prehistoric populations
In hunter-gatherer communities these conditions are not
met. Hunter-gatherer communities are characterised by
little economic differentiation, and resources transferred
to children (e.g. knowledge, social relations) are of the
kind that has very limited impact on social position – and
hence reproductive success – of the offspring14. A similarly
low transmission and variability of resources is observed
in populations of horticulturalists15. However, polygyny
may nevertheless spread in these groups as a male repro-
ductive strategy. The key factor is the level of male con-
tribution to offspring maintenance16. In the communities
in which women’s activities are prevalent in food acquisi-
tion (especially among gatherers), men can afford numer-
14 Smith et al. 2010b; Smith et al. 2010a, 85–93.
15 Gurven et al. 2010.
16 Marlowe 2003, 293–302; 2005, 56; Fortunato/Archetti 2009,
152–153.
ous offspring without bearing additional costs connected
with gaining resources. In such situations, a man’s mating
success will translate into his reproductive success. At the
same time, the “voice” of women may become more im-
portant17, and for them a criterion for choosing partners
will be physical fitness rather than resources. This is con-
firmed by a distinct correlation between the areas of high
pathogen stress (tropics in particular) and increased rates
of polygyny18.
Unlike among hunters-gatherers, resources in far-
ming and pastoral populations (animal herds, farmland)
can be predictably controlled and passed on from gener-
ation to generation19. Therefore, the “vertical” trans-
mission of resources plays a crucial role in the survival
and reproductive success of offspring. Consequently,
groups of animal herders and farmers show significant
differences with regard to the wealth at the disposal of in-
dividuals or lineages20. A high level of economic differ-
entiation may create evolutionary pressure, activating the
polygyny threshold model, although empirical research
on contemporary pre-modern farming21 and herding22 so-
cieties gave ambiguous results in this respect. Wealth
variability may also result in higher reproductive benefits,
obtained in rich families by investing in the marriage of
sons (bride price), and in poor families by investing in
daughters, who could “marry up”23. In farming/herding
populations, there are also other factors which may favour
monogamy or polygamy. The most important seems the
depreciation of resources resulting from sharing them
among offspring24. In herding groups, inherited resources
are animal herds which can be quickly reproduced. There-
fore, in subsequent generations, the value of transmitted
goods per family member should not vary significantly,
even provided that a kinship group grows steadily. What
is more, the specific character of resources available to
herdsmen allows using them not only as a subsistence
basis, but also as a means of building social relations25.
In the case of farming communities the situation is
different. Here, basic heritable resources are land use
rights, which increase their value with the intensification
17 Low 2005, 68.
18 Marlowe 2003, 293–298; Ember/Ember/Low 2007, 431–437.
19 Shennan 2011, 920–921.
20 Borgerhoff Mulder et al. 2010; Shenk et al. 2010; Smith et al.
2010a, 85–93.
21 Borgerhoff Mulder 1990; Strassmann 2000.
22 Sellen/Borgerhoff Mulder/Sieff 2000.
23 Hartung 1982; Roth 2004, 73–74.
24 Fortunato/Archetti 2010, 154.
25 E.g. Shennan 2011, 921.
Fig. 1: Tombstone of Count von Gleichen and two accompanyingwomen. The legend refers it to Ernst von Gleichen (1160–ca. 1246)who supposedly obtained the pope’s permission to have two wivesafter having returned from a crusade. Erfurt cathedral, 2nd half of the13th century (photo Marzena Przybyła)
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Marcin S. Przybyła, Mating systems in prehistoric populations 211
of farming26 or when the possibility of extending the acre-
age becomes limited. Under such conditions, dividing
resources (land) between numerous offspring will result
in depreciation of their value per family member in suc-
cessive generations. To prevent this, property in farming
societies is inherited by only one child, usually the oldest
or the youngest son27, or a tendency to limit the number of
male offspring can appear28. Such tendencies may pro-
mote monogamy as a prevailing mating system.
The above review leads to the conclusion that the
presence or domination of a specific mating system does
not stem from a single determinant, but from the combi-
nation of various factors. Three of these can be mentioned
as the most important: (i) level of male investment in sur-
vival and reproductive success of offspring, (ii) differenti-
ation of the value of resources at the disposal of men and
(iii) extent to which resources are permanently depreci-
ated when divided among offspring (Fig. 2). Polygyny in a
hypothetical population will be promoted by a low value
26 Adler 1996.
27 E.g. Boone 1986, 868–869; Chu 1991.
28 E.g. Beise/Voland 2002.
of men’s contribution, combined with a low level of econ-
omic differentiation and its low influence on the offspring’s
social position. Such a situation occurs in numerous con-
temporary hunter-gatherer communities. However, high
values of these factors may create conditions described in
the polygyny threshold model. The level that the econ-
omic inequality and parental investment in their off-
spring’s social position must reach to make polygyny a fa-
voured strategy will then depend on the degree to which it
is women who choose their partners. It seems, however,
that the patriarchal suzerainty over the female part of the
population, especially with respect to marriage, can be
viewed as one of the predictors of polygyny29. On the other
hand, an important factor reducing the impact of econ-
omic inequality between men on selecting polygyny will
be the extent to which resources controlled by men lose
their value after they are divided among offspring. There-
fore, polygyny is most likely to appear in populations en-
gaged in animal husbandry (both mobile pastoral groups
and settled herders), especially if they show distinct econ-
omic differentiation. Monogamy will be preferred in far-
29 E.g. Marlowe 2003, 284; Low 2005, 68–69.
Fig. 2: Factors promoting polygyny (dark area) and monogamy (light area)
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212 Marcin S. Przybyła, Mating systems in prehistoric populations
ming groups occupying densely populated centres of
settlement oekumenes. However, its competitiveness with
respect to polygyny will decrease in marginal zones (with
enough room for economic expansion) and in groups with
more distinct economic differentiation due to availabil-
ity of resources other than cultivated land (supplemen-
tary role of animal husbandry, exploitation of rare re-
sources).
The aim of the present paper is to confront the above
regularities with the archaeological record. The study de-
scribed below is based on the analysis of grave goods from
selected Neolithic, Bronze Age and Early Iron Age cem-
eteries in Central Europe. It attempts to identify differ-
ences in male and female grave assemblages, which might
be interpreted as resulting from polygyny or monogamy,
and to then compare them with parameters describing
economic differentiation and the production system of the
discussed populations.
Sources and methodology
Criteria of source selection
The investigation is based on data obtained from twelve
cemeteries. The sites were selected using the criterion of
the number of graves with reliably determined age and sex
of the deceased. Selected for analysis were the graves of
adults (juvenis/adultus-senilis) with no traces of looting or
other secondary actions which might have influenced the
homogeneity of the contents. Burials where the unam-
biguous determination of sex was impossible due to the
small number of bones (in cremations) or their poor state
of preservation were not included30. The selected cem-
eteries represent a broad span of time. The cemetery at Tis-
zapolgár-Basatanya, kom. Hajdú-Bihar31 comes from the
early Eneolithic of the Carpathian Basin (the Tiszapolgár
culture), while the site at Budakalász, kom. Pest32 dates to
the end of this period (the Baden complex). The Early
Bronze Age is represented by the cemetery at Singen,
Kr. Konstanz33 in the Alpine zone, and by roughly contem-
porary sites of the so-called Epi-corded Carpathian Cul-
tural Circle from Iwanowice, pow. Miechów34, Jelåovce,
30 E.g. Kunter 1993, 208.
31 Bognár-Kutzián 1963.
32 Bondár/Raczky 2009.
33 Krause 1988.
34 Kadrow/Machnik/Machnik 1992.
okr. Nitra35 and Branb, okr. Nitra36. The Tumulus Circle
cemetery at Tápé, kom. Szeged is slightly younger37. Sites
such as Bachórz-Chodorówka, pow. Rzeszów38, Pysznica,
pow. Stalowa Wola39, Przeczyce, pow. Zawiercie40 and
Obereching, Kr. Salzburg41 represent local variants of the
Urnfield complex from the Late Bronze Age and the begin-
nings of the Iron Age. Finally, the cemetery at Chotín, okr.
Komárno42 belongs to the so-called Traco-Scythian groups
of the Iron Age in the Carpathian Basin. The broad chro-
nological and spatial span represented by the sites se-
lected for analysis was chosen to reduce the impact of
similarities or differences resulting from historical con-
nections, diffusion or the transmission of cultural traits.
A similar solution to this methodological problem is ap-
plied in comparative studies in anthropology43.
Determination of status on the basis of graveassemblages
The way in which the organisation of social life translates
into burial rites belongs to the most frequently discussed
issues in archaeology44. According to some scholars, the
burial rite has been created for specific social purposes,
thus it reflects an ideal society rather than real relations
within it45. At the same time, in pre-modern communities,
wealth need not translate directly into status within a
group46. Bearing these restrictions in mind, one should
however not forget that funerary evidence remains the pri-
mary, and sometimes the only, source for studying the or-
ganisation of prehistoric societies. Furthermore, for the
issue analysed in this paper, it is important to identify the
economic position of the deceased (which may be more
directly manifested in lavish grave goods), rather than
their actual status within the group.
The evaluation of the wealth of a grave assemblage
can be based on a grading scale established for particular
35 Bátora 2000.
36 Vladár 1973.
37 Trogmayer 1975.
38 Gedl 1994; Szybowicz 1995.
39 Czopek 2001.
40 Szydłowska 1968; 1972.
41 Höglinger 1993.
42 Duåek 1966.
43 Compare: Mace/Holden 2005, 119–120; Murdock/White 2006.
44 For a review of older literature, see: Bartel 1982; Ames 2008,
495–501.
45 Bernbeck 1997, 264–268, and literature cited in this work.
46 E.g. Müller/Bernbeck 1996, 1–2; 16–19.
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Marcin S. Przybyła, Mating systems in prehistoric populations 213
elements, either a priori47 or relatively48. Another method
consists in a simple count of artefacts or of their types, ne-
glecting differences in their value49. Due to the cultural
diversity of the sources, in the present study this latter,
simpler solution was used. Grave wealth was evaluated
proceeding from the number of artefacts discovered, with
additional division into ceramic and non-ceramic equip-
ment. To exclude the potential impact of differences in
male and female costume, the calculations of wealth stan-
dards were conducted for each sex separately. Addition-
ally, in order to control the results obtained, for some cem-
eteries a more sophisticated, relative method of wealth
estimation was also used (the so-called “scarcity index”)50.
The picture it provided is comparable with that obtained
by the simple counting of inventory units (e.g. Fig. 3).
47 E.g. according to the criterion of the time needed to manufacture
it – Peregrine 1991.
48 Hodson 1977, 406–410; Jørgensen 1988, 21–22.
49 Hedeager 1992, 103.
50 Hodson 1977, 406–410; Jørgensen 1988, 21–22.
Sex-biased asymmetry of wealth distribution(WDA)
Preliminary investigations of differences in wealth of
grave goods showed that in spite of the very simple
method used to evaluate grave wealth, the result was
close to expectations: the number of graves in successive,
arbitrarily established wealth classes decreased, resulting
in a kind of socio-economic “pyramid” (Fig. 4). The only
exception was the cemetery at Singen, where the majority
of graves belonged to the richest categories. At this stage
of the investigation, ceramic and non-ceramic equipment
was treated separately. However, in further studies these
two categories were analysed together, since in some of
the cemeteries vessels were not deposited in the graves at
all, and in the others their number was distinctly corre-
lated with the number of other elements of equipment.
The initial stage of research brought one more obser-
vation. Although the percentage of male and female
burials in particular wealth categories was similar in the
majority of cemeteries, some sites showed an interesting
asymmetry. There, a higher share of female graves at-
tributed to the highest categories was accompanied by a
Fig. 3: Evaluation of graves wealth obtained by using two different methods:older and younger phase of the Early Bronze Age cemetery in Iwanowice (Małopolska)
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214 Marcin S. Przybyła, Mating systems in prehistoric populations
Fig. 4: Percentage of female and male graves within arbitrarily assumed equipment classes: Bachórz-Chodorówka (1), Branb (2), Budakalász(3), Chotín (4), Iwanowice (5), Jelåovce (6), Obereching (7), Przeczyce (8), Pysznica (9), Singen (10), Tápé (11), Tiszapolgár-Basatanya (12)
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Marcin S. Przybyła, Mating systems in prehistoric populations 215
high frequency of male graves totally devoid of grave
goods. The overrepresentation of high-status female
graves and a high number of very modest male burials
may be regarded as manifesting a higher “social mobility”
of women. In the populations characterised by high social
differentiation, men who gain control over available re-
sources become objects of rivalry among women, includ-
ing women from lower classes. This may result in a ten-
dency for women to seek social advancement through
marriage, and the reproductive success of a female be-
comes dependent on the economic status of her husband,
rather than that of her parents51.
On the one hand, in such a situation the polygyny
threshold may operate, and on the other it may strengthen
patriarchal control over the marital decisions of women,
reducing the competitiveness of monogamy52. Asymmetry
in wealth distribution between male and female burials
may thus be perceived as an archaeological indicator for a
high probability of polygyny. A similar interpretation of
this phenomenon has already been proposed in studies on
selected prehistoric cemeteries53.
In order to perform further analyses, the observed
asymmetry of wealth distribution had to be expressed
quantitatively. This was achieved using the following for-
mula:
WDA =Fw : (Fw + Fp)
Mw : (Mw + Mp)
where: Fw – the number of female graves in which the
number of artefacts is equal to the value in the third quar-
tile of the class of female graves (sumptuous female
graves), Fp – the number of female graves devoid of grave
goods (poor female graves), Mw – the number of male
graves in which the number of artefacts is equal to the
value in the third quartile of the class of male graves
(sumptuous male graves), Mp – the number of male graves
devoid of grave goods (poor male graves).
In the calculations, a separate wealth classification was
assumed for male and female graves in order to make up
for possible differences in the standards of equipment re-
sulting from differences in male and female costumes and
burial rituals. A calculated value equal to 1 indicates a
symmetric distribution of wealthy and poor graves of men
and women. A value higher than 1 means that the share of
the poorest burials is higher for men than for women, or
51 E.g. Boone 1986; Shennan 2002, 181–197.
52 Compare: Low 2005, 68–68.
53 Shennan 1975, 285–286; Ter}an 1992, 81.
that the percentage of the most lavishly equipped graves is
higher for women, or both.
Economic inequality
Economic inequality belongs to the factors which may in-
fluence the selection between the two competing mating
systems. Variability of grave wealth, strictly connected
with economic inequality, was determined using the Gini
coefficient – a measure of the inequality of a distribution,
based on the Lorenz curve. The Gini coefficient, long ap-
plied in econometrics, has for some time also been used in
anthropological cross-cultural studies54 and in archaeol-
ogy55. Its value can range from 0 (equal distribution – tot-
ally egalitarian society) to 1; contemporary societies with a
monetary economy are characterised by values from 0.25
to approximately 0.656. As with the wealth distribution
asymmetry (WDA) coefficient, in analyses using the Gini
coefficient grave wealth was evaluated based on the total
number of artefacts in the assemblage. These analyses
were performed for male burials only – which results from
the assumption that in highly hierarchical societies, the
social status of a man is inherited (contrary to the more
“mobile” status of a woman). Thus, male graves may re-
flect a “rigid” picture of unequal access to resources.
Production system – attribution basedon assignment to cultural units
Another factor which may promote one of the two main
mating systems is the predominant form of subsistence.
In deciding what production system should be assigned to
the analysed sites, one should choose between intensive
farming and various forms of economy relying on animal
herding (stationary herding-farming economy, transhu-
mance, mobile pastoralism). The “farming” or “herding”
attribution may be chosen according to economic strategy
thought to be characteristic of the cultural tradition repre-
sented by a given site. Such an approach takes into ac-
count a broad spectrum of observations, but its disadvan-
tage lies in losing a local perspective.
According to the predominant view among scholars,
a significant role of animal herding – or even a pastoral
lifestyle – should be accepted for the population using the
54 Bowles/Smith/Borgerhoff Mulder 2010, 11.
55 Ames 2008, 499–502.
56 Smith et al. 2010b, 30.
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216 Marcin S. Przybyła, Mating systems in prehistoric populations
cemetery at Tiszapolgár-Basatanya57. This view has been
confirmed by studies on local environmental changes in
the Tisza basin, which revealed distinct traces of defore-
station and animal grazing dated to the mid-fifth millen-
nium BC58. Herding (with the possible local occurrence of
intensive agriculture) is also taken into consideration for
groups representing the Epi-corded Carpathian Cultural
Circle (cemeteries at Branb, Iwanowice and Jelåovce)59 and
for the Tumulus complex (cemetery at Tápé). With regard
to this latter tradition, the old hypothesis about the parti-
ally mobile lifestyle of its populations remains valid60. Fi-
nally, a herding economy and a semi-nomadic or even no-
madic lifestyle is seen as characteristic of the so-called
Vekerzug culture, represented by the cemetery at Chotín61.
However, recent studies have indicated that in the Early
Iron Age the Great Hungarian Plain was also inhabited by
farming groups62.
The predominant form of the economy, or even the
mobility of the Baden complex populations from north-
western Hungary (cemetery at Budakalász) are still dis-
cussed in the literature, although palaeobotanic studies
allow us to assume an important role of farming in this
case63. Permanent settlement, as well as the cultivation of
cereals and leguminous plants, are well documented for
sites in southern Germany, both dating to the Early (cem-
etery at Singen) and Late (cemetery at Obereching) Bronze
Age64. A mixed farming-herding economy is suggested for
the Lusatian culture in the Oder and Vistula basins, repre-
sented in the present work by the cemeteries at Bachórz-
Chodorówka, Przeczyce and Pysznica. Regional studies
provide strong evidence for intensive farming65, while
studies on herd structure indicate a high percentage of
pig, typical of settled populations66.
Grading scale for non-agricultural resources
An examination of specific environmental factors and cer-
tain elements of the archaeological record for each site indi-
vidually may provide us with a more objective method of
57 E.g. Sherratt 1981.
58 E.g. Sümegi 2005, 127; 136–137.
59 Kadrow 1996, 68–76.
60 E.g. Kristiansen 1998, 414.
61 Chochorowski 1985, 133–134.
62 Chapman/Magyari/Gaydarska 2009.
63 Patay/Herbich/Sümegi 2008, 91–94; Gyulai 2010, 89–92; compare
e.g. Whittle 1996, 123; Bondár/Raczky 2009, 480.
64 E.g. Primas 2008, 20–21; 88–89.
65 E.g. Urban 2009.
66 Harding 2000, 134; 140.
evaluating how dependent the populations in question
were on farming. The proposed “grading scale for non-agri-
cultural resources” uses the following variables (ranging
from 0 to 2): vegetation zone where a given site is located
(high score for geographical conditions unfavourable for
farming), importance of animals in the burial rite (high
value of this variable indicates a socio-ritual role of animals,
which is typical of herding groups regardless of its symbolic
interpretations67), access to rare resources (actually ex-
ploited during the period in question) and traces of partici-
pation in long-distance exchange (the participation in long-
distance exchange and control over rare resources might
additionally increase social differentiation and provide
heritable resources other than arable land). A total value of
all the variables produces a coefficient, which may be com-
pared with the WDA index for particular sites (Tab. 1).
The attribution to a vegetation zone was based on gen-
eralised isopollen maps68 representing two chronological
horizons: ca 2000 BC (for sites dating to the Eneolithic and
earlier parts of the Bronze Age) and ca 0 BC/AD (for sites
from the later Bronze Age and the Early Iron Age) (Fig. 5).
The highest score was attributed to the steppe zone and a
slightly lower one to the zone of sub-mountainous veg-
etation. As for the variable describing the share of animal
sacrifices in the burial rite, the highest score was attributed
to the occurrence of animal graves or to a high (over 20%)
percentage of graves containing the remains of sacrificed
animal carcasses69. In all other cases where animal remains
were found in graves, this variable had the value of 170. Ac-
cess to rare resources was granted the highest score in the
case of two Alpine sites, located in the area which was the
economic background for “mountainous” populations en-
gaged in the most intensive exploitation of copper deposits
in prehistoric Europe as a whole71. A slightly lower score
was attributed to the cemeteries at Jelåovce and Branb, lo-
cated close to the zone of the very early exploitation of
copper ore in central Slovakia72, and to the cemetery at Tis-
zapolgár-Basatanya, adjacent to the region where native
copper had already been extracted in the Eneolithic73. The
last variable encompassed the cases where objects of defi-
nitely exotic provenience were recorded. This category in-
67 Stadler 2010.
68 Huntley/Birks 1983.
69 Bognár-Kutzián 1963, 377–386; Duåek 1966, 17–24; Vladár 1973,
165; Bátora 2000, 459–460.
70 Kubasiewicz 1972; Trogmayer 1975, 149; Kadrow/Machnik/Mach-
nik 1992; Höglinger 1993, 53–54; Szybowicz 1995, 20–21; Bátora 2000,
459–460; Czopek 2001, 164–166; Blischke 2002, 111–112; Gál 2009.
71 E.g. Krause 1988, 214–245; Shennan 1995, 300–308.
72 E.g. Schreiner et al. 2005.
73 E.g. Whittle 1996:117–120.
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Marcin S. Przybyła, Mating systems in prehistoric populations 217
Tab. 1: Data for calculating “non-agricultural resources” score
Site Vegetation zone Animals in burial rite Access to rare resources Access to exotic goods
Bachórz-Chodorówka
Mixed-deciduous forest –0 point
Charred bone remains ingraves, two gravescontaining only the animalbones (including pigremains) – 2 pts
None – 0 pts Only local products – 0 pts
Branb Montane mixeddeciduous-conifer forest –1 point
Numerous graves (37 %)with cattle remains – 2 pts
Neighbourhood of apotential zone of copperacquisition in centralSlovakia – 1 pts
Few faience beads and shellsof Mediterranean origin – 1 pts
Budakalász Montane mixeddeciduous-conifer forest –1 point
Scarce (3 %) gravescontaining the remains ofanimals (most often wildanimals) – 1 pts
None – 0 pts Only local products – 0 pts
Chotín Steppe – 2 points Few horse graves, horsesdeposited in human gravesas well – 2 pts
None – 0 pts Graves with amber beads andsingle gold artefacts andmirrors – 1 pts
Iwanowice Mixed-deciduous forest –0 point
Few animal bones in gravepits – 1 pts
None – 0 pts Few faience beads – 1 pts
Jelåovce Montane mixeddeciduous-conifer forest –1 point
Single graves with animalbones (cattle) – 1 pts
Neighbourhood of apotential zone of copperacquisition in centralSlovakia – 1 pts
Few faience beads – 1 pts
Obereching Montane mixeddeciduous-conifer forest –1 point
Several graves (7.6 %) witpig or sheep remains – 1pts
Direct background of themain zone of copperacquisition – 2 pts
Only local products – 0 pts
Przeczyce Mixed-deciduous forest –0 point
20 graves with cattle andsheep remains – 1 pts
None – 0 pts Only local products – 0 pts
Pysznica Mixed-deciduous forest –0 point
Charred bone remains ingraves (small ruminants inparticular) – 1 pts
None – 0 pts Only local products – 0 pts
Singen Montane mixeddeciduous-conifer forest –1 point
No data available – 0 pts Direct background of themain zone of copperacquisition – 2 pts
Daggers from the “Atlanticmetallurgic centre” – 2 pts
Tápé Steppe – 2 points Pits with animal andhuman bones found oncemetery, few graves withanimal bones – 1 pts
None – 0 pts Only local products – 0 pts
Tiszapolgár-Basatany a
Steppe – 2 points Fragments of animalcarcasses in almost half ofgraves (mainly cattle, butalso goat, sheep and pig).Moreover, dog burialsaccompanying humanburials – 2 pts
Potential neighbourhoodof native copper depositsin NE Hungary – 1 pts
Both stone and copperartefacts (few) may be of alocal origin – 0 pts
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218 Marcin S. Przybyła, Mating systems in prehistoric populations
Fig. 5: Localisation of the investigated sites against the background of main vegetation zonesaround 2000 BC and 0 BC/AD (after Huntley/Birks 1983): Bachórz-Chodorówka (Ba), Branb (Br),Budakalász (Bu), Chotín (Cho), Iwanowice (Iw), Jelåovce (Je), Obereching (Ob), Przeczyce (Pr),Pysznica (Py), Singen (Si), Tápé (Ta), Tiszapolgár-Basatanya (Ti)
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Marcin S. Przybyła, Mating systems in prehistoric populations 219
cluded faience beads74 and luxury products of the Iron Age
nomadic peoples75. The highest score was attributed to the
collection of “Atlantic” daggers from the Singen cemetery,
whose foreign provenience – determined based on stylistic
assessment – was confirmed by metallographic analyses76.
ResultsNumerical values obtained in the analysis (Tab. 2) were
listed and compared (Fig. 6). The comparison took into
consideration some additional attributes, such as site
chronology and sex ratio in the analysed populations, ex-
pressed as percentage of female graves.
Examination of grave wealth inequality revealed high
differentiation within the analysed sample. The Gini co-
efficient oscillated between a very low value for the cem-
etery at Bachórz-Chodorówka (0.23) and high results (ex-
ceeding 0.70) for Branb, Jelåovce, Iwanowice and Tápé,
while the mean value for the whole sample amounted to
0.58. A slight correlation can be noted between chrono-
logy and the Gini coefficient value (Fig. 6:A). Dispropor-
tion in grave equipment decreases on sites from later seg-
ments of the Bronze Age and from the Early Iron Age.
Another, although not very significant correlation can be
shown between the Gini coefficient value and the “wealth
74 In spite of the fact that their Mediterranean origin remains open to
question – e.g. Harding/Warren 1973.
75 Duåek 1966, 35–36; Vladár 1973, 152–154; Kadrow/Machnik/Mach-
nik 1992, 81; Bátora 2000, 340–342.
76 Krause 1988, 56–63.
distribution asymmetry” index (Fig. 6:B). The majority of
cemeteries with a WDA value higher than 1 (i.e. cemeteries
used by communities where polygyny could potentially
have occurred) also has a high Gini coefficient value (over
0.65). It is worth noting that this group comprises sites at-
tributed to herding or even pastoral cultural traditions
(black signatures on the diagram). It should also be men-
tioned that the two outliers – the cemeteries at Obereching
and Singen – are distinguished by a low number of graves,
a particularly high standard of grave goods and a low per-
centage of female burials (Tab. 2, Fig. 4).
Another diagram (Fig. 6:C) compares the proportion
of female burials with the “wealth distribution asym-
metry” index. Half of the investigated sites fall into the
range of approximately equal sex distribution (40–50 % of
graves are female), but at the same time show a strong dif-
ferentiation of the WDA index. A slight overrepresentation
of male graves at these sites may result from the applied
methods of sex determination77. However, it is sympto-
matic that three cemeteries with more than 50 % female
graves also have a low WDA index, while three cemeteries
where this index is high (Iwanowice, Obereching and
Singen) show a distinct overrepresentation of male
burials.
The most significant correlation is found when com-
paring the “non-agricultural resources” score with the
WDA index (Fig. 6D). For the five cemeteries with a low
score, which suggests strong dependence on agricultural
resources, the value of the WDA index falls within the
77 Compare: Weiss 1972; Volk/Büchner 1997, 27–29; Chamberlain
2009, 90–91.
Tab. 2: Basic numerical data describing the analysed sites
Site Chronology Number ofana-lysed graves
Share offemales burials (%)
Gini index formales burials
WDA index “non-agriculturalresources” score
Bachórz-ChodorówkaPhase A-AB
14–11 c. BC 51 60 0.23 0.85 2
Branb 21–18 c. BC 137 48 0.70 1.72 5Budakalász 32–25 c. BC 113 62 0.64 0.90 2Chotín 6–5 c. BC 116 69 0.50 1.00 5Iwanowice 21–17 c. BC 90 31 0.80 1.27 2Jelåovce (Nitra Group) 21–18 c. BC 122 40 0.72 1.32 4Obereching 11–10 c. BC 49 30 0.42 1.22 4Przeczyce 10–8 c. BC 96 46 0.44 0.76 1Pysznica 8–4 c. BC 166 40 0.64 0.69 1Singen 22–20 c. BC 33 30 0.46 1.66 5Tápé 17–15 c. BC 335 46 0.75 1.16 3
Tiszapolgár-Basatanya 45–39 c. BC 99 47 0.67 1.19 5
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220 Marcin S. Przybyła, Mating systems in prehistoric populations
range from 0.69 to 1.27, with a mean of 0.89. For the re-
mainder of the sites, where “non-agricultural resources”
scored at least 3, the mean WDA index was as high as 1.32.
DiscussionIn the present study, overrepresentation of female burials
in the richest equipment classes, accompanied by over-
representation of men in poor burials, is regarded as an in-
dicator of polygyny. However, an alternative explanation
of the high WDA index may be serial monogamy, that is a
situation in which a man has several wives over his whole
life, but only one in a given time span. The most probable
reason for serial monogamy is a higher mortality rate
among women (particularly young women) than men. In
this case we should expect that an overrepresentation of
female graves among richer classes of burials will corre-
late with a significantly lower mean age of the deceased
women compared to the men within the whole popu-
lation, and within the group of wealthier burials in par-
Fig. 6: Comparison of results. In diagram B light signatures stand for sites with farming attribution, black signatures stand forherding attribution. Abbreviations of site names: Bachórz-Chodorówka (Ba), Branb (Br), Budakalász (Bu), Chotín (Cho),Iwanowice (Iw), Jelåovce (Je), Obereching (Ob), Przeczyce (Pr), Pysznica (Py), Singen (Si), Tápé (Ta), Tiszapolgár-Basatanya (Ti)
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Marcin S. Przybyła, Mating systems in prehistoric populations 221
ticular. In contrast, the difference between the mean age
of women and men is expected to be less among the popu-
lations with a lower WDA index. However, observations
made for all cemeteries do not match these predictions
(e.g. Fig. 7). The mean age of buried women is usually (but
not always) slightly lower than that of men, but variation
of this factor does not correlate completely with the WDA
index. In consequence, polygyny seems the more prob-
able explanation for the observed pattern.
One of the sites where the asymmetry in male and fe-
male grave assemblages is clearest (WDA index = 1.72) is
the cemetery at Branb. This site was previously thoroughly
analysed by Susan Shennan78. Although Shennan applied
different methods for determining the wealth of the
burials and took into account the precise chronology of
grave assemblages, her conclusions are similar to those
obtained in the present study. Shennan regards the pre-
dominance of female burials among sumptuous graves as
a manifestation of polygyny. Sex determination for
children’s graves (based on archaeological criteria) en-
abled her to notice a distinct rise in the number of lavishly
equipped women’s burials compared to few wealthy girls’
burials, which may be interpreted as a result of women’s
social advancement through marriage79.
In this case, the “social mobility” of women may re-
sult from a large inequality among men with regard to
78 Shennan 1975.
79 Shennan 1975, 285–286.
control of resources. The Gini coefficient for male inven-
tories from the Branb cemetery amounts to no less than
0.70. If the differentiation in grave wealth really reflects
economic differentiation, we would here be dealing with a
very high degree of economic inequality80. A similarly
high Gini coefficient value (at least 0.65) was also recorded
for the cemeteries at Iwanowice, Jelåovce, Tiszapolgár-
Basatanya and Tápé. At each of these sites one can notice
a high share of wealthy female graves or an overrepresen-
tation of men among burials devoid of grave goods.
The cemetery at Tápé has attracted particular atten-
tion due to a complex of female graves equipped with
pairs of pins and composite neck ornaments. According to
Jobst Blischke81, these burials represent a group of mature
women of outstanding status, whose special position
would also be also manifested by the location of their
graves in two isolated zones of the cemetery. It is worth
noting that Blischke’s social interpretation of the Tápé
cemetery differs completely from the interpretation pro-
posed in this paper. In Blischke’s opinion, the remarkable
equipment might reflect the women’s high, cult-related,
social position and the weakening of the “men’s world”,
which suffered a global crisis82. In contrast, the evolution-
istic perspective adopted in this paper suggests a general
correlation between the manifestation of wealth in female
80 Compare: Bowles/Smith/Borgerhoff Mulder 2010, 11.
81 Blischke 2002, 59–75; 153.
82 Blischke 2002, 272.
Fig. 7: Age of women and men buried on two cemeteries with the lowest WDA index, and two with the higher one (the cemeteries at Branband Singen having the highest WDA index have been not taken into account since they did not provide longer series of burials with preciseage determination). As “richer” graves are defined those with the number of artefacts larger then the mean for the whole cemetery
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222 Marcin S. Przybyła, Mating systems in prehistoric populations
costume and the domination of men in real social re-
lations83.
In communities characterised by a productive econ-
omy and inheritance through the male line, increased
economic inequality among men should intensify the
competition among women for the most affluent partners.
This stems from the rule quoted above, which posits that
in such populations a woman’s reproductive success is
more strongly influenced by the social status of her hus-
band than that of her parents. From such a perspective,
lavishness of female dress (reflected in burial) might be
seen as a sign of parental investment in the form of dowry
that increased their daughter’s chances to find a proper
partner or even to marry up the social ladder. The institu-
tion of dowry exists mainly in monogamous populations,
where the competition for the most affluent men is par-
ticularly intensive84. However, one should remember that
in communities where the status of a woman depends on
the status of her husband, it is much more likely that fe-
male costume is used to manifest the value of male-con-
trolled resources. With prestige rivalry, this may lead to a
higher degree of refinement of the shapes, sizes and dec-
oration of female ornaments. A good example seems to be
represented by the Tumulus Circle cemeteries (17th–14th
century BC), such as the above-mentioned site at Tápé
(Gini coefficient = 0.75, WDA index = 1.16). In this cultural
tradition, a remarkable lavishness of female graves is
commonly recorded. There is also a tendency to crowd to-
gether the motifs on bronze ornaments and to enlarge
some of them – bronze pins in particular – to the limits of
their functionality85. Thus, although the form and diver-
sity of female dress in this phase of the Bronze Age might
have served behaviours related to social communi-
cation86, one cannot exclude that the tendency towards
the refinement of material culture itself was a process
initiated by sexual selection.
In populations with little economic inequality among
men, a permanent competition for mating partners is
more likely to take place within the male part of a popu-
lation87. This can be illustrated by the cemetery at
Przeczyce, characterised by a relatively low value of the
Gini coefficient (0.44) and a very low WDA index (0.76).
The distribution of wealth at the site revealed a distinct
bias towards male graves. However, this need not neces-
sarily reflect patriarchal dominance alone – as suggested
83 See Shennan 2002, 205.
84 Compare: Gaulin/Boster 1991; Roth 2004, 74–78.
85 E.g. Wiegel 1994; Innerhofer 2000.
86 E.g. Sørensen 1997.
87 Shennan 2002, 196–197.
by the investigator of the site88. Instead, this may reflect
the tendency towards a higher degree of refinement in
manifesting wealth by men, i.e. the development (under
conditions of competition for resources) of material mani-
festations of prestige, or even of some “epigamous traits”
connected with male rivalry for access to female partners89.
In this case, too, the distinctly higher wealth of the de-
ceased of one of the sexes does not translate into their
higher rank in a society.
Substantial economic inequality among men and the
resulting higher “social mobility” of women do not auto-
matically mean that polygyny will be selected. The com-
petitiveness of polygamy will be reduced by the fact that
the offspring of successive wives are likely to receive less
and less resources from men and by the impossibility to
share durable resources among offspring. Additional,
non-agricultural resources may diminish the importance
of the first factor. Such a situation is recorded for the cem-
eteries at Obereching and Singen (WDA index: 1.22 and
1.66, respectively). In both cases we should take into ac-
count that local populations participated both in supply-
ing provisions to the centres of copper ore mining and in
redistributing their products. The low differentiation of
grave assemblages on both sites (Gini coefficient below
0.50) may be explained by their generally high wealth.
However, it cannot be excluded that this result was af-
fected by the low number of female graves at the two sites.
The constraint on the spread of polygyny related to
the depreciation of durable resources after dividing them
among offspring may be less important for populations
which rely primarily on animal herding. Among the ana-
lysed sites, a “culturally” ascribed herding attribution ac-
companied by a high “non-agricultural resources” score is
recorded for the Eneolithic cemetery at Tiszapolgár-Basat-
anya, the sites at Branb and Jelåovce connected with the
Epi-corded complex, the Tumulus culture cemetery at
Tápé and the Iron Age Vekerzug culture cemetery at Cho-
tín. All these sites had a WDA index equal to at least 1. Per-
haps it is no coincidence that the WDA value was the lo-
west in the case of the cemetery at Chotin, which – as the
only one in the group – also has a rather low coefficient of
grave good inequality among males (0.50). We may make
a cautious suggestion that polygamous marriages were
admissible in the populations using these cemeteries. Per-
haps the site at Iwanowice should also be added to this
group. Undoubtedly, it represents a cultural tradition
characterised by a production system which relied – at
88 Szydłowska 1972, 201.
89 Compare e.g. Boone 1998; Bliege Bird/Smith 2005.
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Marcin S. Przybyła, Mating systems in prehistoric populations 223
least at the beginning of its development – on mobile
herding90. On the other hand, the low “non-agricultural
resources” value (2 pts) for this cemetery corresponds for
this cemetery with the discovery of a neighbouring, large,
long-lasting settlement with numerous pits used to store
farm produce91. Regardless of the conclusions about the
economy of the Iwanowice population, it is worthwhile
noting that the high WDA index (1.27) correlates with
the highest recorded coefficient of grave good inequality
among male graves (0.80).
ConclusionsGeneralising the above observations, one can state that
the results of the present study agree with the previously
formulated expectations. The elements of the archaeologi-
cal record which may indicate the occurrence of polygyny
generally correlate with two factors which favour it: high
economic inequality among males and an economy in
which resources are not depreciated when shared among
numerous offspring (thus not decreasing their fitness). A
second general conclusion is methodological. It seems
that the results presented here allow us to accept that ar-
chaeological sources can be analysed following the pro-
cedures typical of anthropological cross-cultural studies,
that is by comparing selected variables which describe un-
connected populations92. Further development of this re-
search procedure may, on the one hand, enrich studies on
the problem of how the actual functioning of prehistoric
populations is reflected in the archaeological record and
provide information about particular communities, while
on the other hand it may elucidate the development over a
long time period of more universal phenomena studied by
the ecology of human behaviours.
AcknowledgementsA preliminary version of this paper was presented in 2010
at the EAA Annual Meeting in The Hague, as part of the
session dedicated to women’s status in prehistory. I
warmly thank the organisers – Bettina Arnold and Hrvoje
Potrebica – for enabling me to present the results of my
studies. The paper in its present form owes much to the
90 Kadrow 1996:68–76.
91 Kadrow 1991.
92 As it has already been postulated before – eg. Peregrine 2001.
critical remarks and bibliographic suggestions of Stephen
Shennan, Peter Bogucki and an anonymous reviewer, for
which I am also very grateful. I would like to thank Mag-
dalena Skoneczna for discussions that resulted in the
method used to evaluate the subsistence of the popu-
lations investigated here. The translation into English was
carried out by Piotr Godlewski. The paper took its final
shape – in respect of both its merits and form – thanks
to critical and inspiring comments of Marzena Przybyła,
whom I wish to thank very much for her help and non-
schematic approach to the issues presented here.
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